Professor Charles Wellman - Publications
A novel reproductive strategy in an Early Devonian plant. Current Biology, 30(9), R388-R389.
Bernard Owens (1938–2019). Palynology, 44(2), 195-204.
Mid-Devonian Archaeopteris roots signal revolutionary change in earliest fossil forests. Current Biology, 30(3), 421-431.e2. View this article in WRRO
Professor Bernard Owens (1938–2019). Review of Palaeobotany and Palynology, 276, 104184-104184.
Wall ultrastructure of the Permian pollen grain Lueckisporites virkkiae Potonié et Klaus 1954 emend. Clarke : Evidence for botanical affinity. Review of Palaeobotany and Palynology. View this article in WRRO
Reconstructing the terrestrial flora and marine plankton of the Middle Devonian of Spain: implications for biotic interchange and palaeogeography. Journal of the Geological Society, 177(2), 315-324.
Filamentous green algae from the Early Devonian Rhynie chert. PalZ. View this article in WRRO
Palynological analysis of Upper Ordovician to Lower Silurian sediments from the Diyarbakir Basin, southeastern Turkey. Review of Palaeobotany and Palynology, 263, 28-46. View this article in WRRO
Edwin George (‘Ted’) Spinner (1938–2018). Palynology, 43(2), 184-187. View this article in WRRO
Palaeontology: The Rhynie Chert Is the Gift that Keeps on Giving. Current Biology, 29(3), r93-r95. View this article in WRRO
An endemic flora of dispersed spores from the Middle Devonian of Iberia. Papers in Palaeontology. View this article in WRRO
Reply to Hedges et al.: Accurate timetrees do indeed require accurate calibrations. Proceedings of the National Academy of Sciences, 115(41), E9512-E9513.
The timescale of early land plant evolution. Proceedings of the National Academy of Sciences of the United States of America, 115(10), E2274-E2283. View this article in WRRO
The Interrelationships of Land Plants and the Nature of the Ancestral Embryophyte. Current Biology, 28(5), 733-745.e2. View this article in WRRO
Palaeoecology and palaeophytogeography of the Rhynie chert plants: Further evidence from integrated analysis of in situ and dispersed spores. Philosophical Transactions of the Royal Society B: Biological Sciences, 373(1739). View this article in WRRO
Ultrastructure and Probable Botanical Affinity of the Enigmatic SporomorphFroelichsporites traverseifrom the Norian (Late Triassic) of North America. International Journal of Plant Sciences, 179(2), 100-114. View this article in WRRO
The classic Lower Devonian plant-bearing deposits of northern New Brunswick, eastern Canada: Dispersed spore taxonomy and biostratigraphy. Review of Palaeobotany and Palynology, 249, 24-49. View this article in WRRO
A key for the identification of cryptospores. Palynology. View this article in WRRO
Dinosaur-plant interactions within a Middle Jurassic ecosystem—palynology of the Burniston Bay dinosaur footprint locality, Yorkshire, UK. Palaeobiodiversity and Palaeoenvironments, 98(1), 139-151. View this article in WRRO
Palynology of the Middle Ordovician Hawaz Formation in the Murzuq Basin, south-west Libya. Palynology, 41(sup1), 31-56. View this article in WRRO
Discussion on ‘Tectonic and environmental controls on Palaeozoic fluvial environments: reassessing the impacts of early land plants on sedimentation’ Journal of the GeologicalSociety, London, https://doi.org/10.1144/jgs2016-063. Journal of the Geological Society, 174(5), 947-950. View this article in WRRO
Middle Ordovician acritarchs and problematic organic-walled microfossils from the Saq-Hanadir transitional beds in the QSIM-801 well, Saudi Arabia. Revue de Micropaléontologie, 60(3), 289-318. View this article in WRRO
New Palynological Investigations in the Paleozoic of Saudi Arabia and Adjoining Areas (Saudi Aramco – CIMP Joint Project). Revue de Micropaléontologie, 60(3), 279-280.
Episodic river flooding events revealed by palynological assemblages in Jurassic deposits of the Brent Group, North Sea. Palaeogeography, Palaeoclimatology, Palaeoecology. View this article in WRRO
A distinctive marginal marine palynological assemblage from the Přídolí of northwestern Saudi Arabia. Revue de Micropaléontologie. View this article in WRRO
Middle Ordovician cryptospores from the Saq-Hanadir transitional beds in the QSIM-801 well, Saudi Arabia. Revue de Micropaléontologie, 60(3), 319-331. View this article in WRRO
Provincial Devonian spores from South China, Saudi Arabia and Australia. Revue de Micropaléontologie. View this article in WRRO
Middle Jurassic vegetation dynamics based on quantitative analysis of spore/pollen assemblages from the Ravenscar Group, North Yorkshire, UK. Palaeontology, 59(2), 305-328. View this article in WRRO
Cryptospores from the Katian (Upper Ordovician) of the Tungus basin: The first evidence for early land plants from the Siberian paleocontinent. Review of Palaeobotany and Palynology, 224, 4-13.
Palaeoecology of a billion-year-old non-marine cyanobacterium from the Torridon Group and Nonesuch Formation. Palaeontology, 59(1), 89-108. View this article in WRRO
A quantitative comparison of dispersed spore/pollen and plant megafossil assemblages from a Middle Jurassic plant bed from Yorkshire, UK. Paleobiology, 41(4), 640-660. View this article in WRRO
Investigating Devonian trees as geo-engineers of past climates: linking palaeosols to palaeobotany and experimental geobiology. Palaeontology, 58(5), 787-801. View this article in WRRO
The terrestrial biota prior to the origin of land plants (embryophytes): a review of the evidence. Palaeontology, 58(4), 601-627.
Morphology and wall ultrastructure of a new and highly distinctive megaspore from the Middle Jurassic of Yorkshire, UK. Review of Palaeobotany and Palynology, 216, 33-43.
Palaeozoic palynostratigraphy of the Arabian plate [a joint project between Saudi Aramco and the Commission Internationale de Microflore du Paléozoïque (CIMP)]. Review of Palaeobotany and Palynology, 212, 1-2.
Late Ordovician–earliest Silurian chitinozoans from the Qusaiba-1 core hole (North Central Saudi Arabia) and their relation to the Hirnantian glaciation. Review of Palaeobotany and Palynology, 212, 60-84.
Spore assemblages from Upper Ordovician and lowermost Silurian sediments recovered from the Qusaiba-1 shallow core hole, Qasim region, central Saudi Arabia. Review of Palaeobotany and Palynology, 212, 111-126.
Conservation ofMale Sterility 2function during spore and pollen wall development supports an evolutionarily early recruitment of a core component in the sporopollenin biosynthetic pathway. New Phytologist, 205(1), 390-401.
Spore assemblages from the Lower Devonian ‘Lower Old Red Sandstone’ deposits of the Northern Highlands of Scotland: the Berriedale Outlier. Earth and Environmental Science Transactions of the Royal Society of Edinburgh, 105(3), 227-238.
Devonian spores from an intra-oceanic volcanic arc, West Junggar (Xinjiang, China) and the palaeogeographical significance of the associated fossil plant beds. REVIEW OF PALAEOBOTANY AND PALYNOLOGY, 206, 10-22.
The nature and evolutionary relationships of the earliest land plants.. New Phytol, 202(1), 1-3.
Lacustrine Nostoc (Nostocales) and associated microbiome generate a new type of modern clotted microbialite. Journal of Phycology. View this article in WRRO
SPORE WALL ULTRASTRUCTURE OF THE LOWER DEVONIAN ZOSTEROPHYLLS RENALIA HUEBERI AND ZOSTEROPHYLLUM DIVARICATUM. INTERNATIONAL JOURNAL OF PLANT SCIENCES, 174(9), 1302-1313.
Reply to Discussion on 'A high-precision U-Pb age constraint on the Rhynie Chert Konservat-Lagerstätte: Time scale and other implications': Journal, 168, 863-872. Journal of the Geological Society, 170(4), 703-706.
Resistance of filamentous chlorophycean, ulvophycean, and xanthophycean algae to acetolysis: Testing Proterozoic and Paleozoic microfossil attributions. International Journal of Plant Sciences, 174(6), 947-957.
Reproductive organs and in situ spores of asteroxylon mackiei kidston & lang, the most complex plant from the lower devonian Rhynie chert. International Journal of Plant Sciences, 174(3), 293-308.
Palaeophytogeography of Ordovician-Silurian land plants. Geological Society Memoir, 38(1), 461-476.
Pennsylvanian (mid-Bolsovian to Asturian) megaspores and large pollen of the Bristol Coalfield, UK. JOURNAL OF MICROPALAEONTOLOGY, 32, 87-106.
402 Pennsylvanian (mid- Bolsovian-Cantabrian; Moscovian) vegetation change in the Bristol Coalfield, UK. 日本花粉学会会誌, 58(Special), 183.
552 How plants conquered the land : An EvoDevo analysis of the spore wall. 日本花粉学会会誌, 58(Special), 249.
ACETOLYSIS AS A SCREEN FOR DEGRADATION-RESISTANT MODERN ALGAL STRUCTURES FOR PALEONTOLOGICAL COMPARISONS. JOURNAL OF PHYCOLOGY, 48, S33-S34.
A timeline for terrestrialization: consequences for the carbon cycle in the Palaeozoic. PHILOSOPHICAL TRANSACTIONS OF THE ROYAL SOCIETY B-BIOLOGICAL SCIENCES, 367(1588), 519-536.
Mid Devonian megaspores from Yunnan and North Xinjiang, China: Their palaeogeographical and palaeoenvironmental significances. Palaeoworld, 21(1), 11-19.
Palaeobotany of the Pennsylvanian (mid-Bolsovian-Cantabrian; Moscovian) Warwickshire Group of the Bristol Coalfield, U.K.: Biostratigraphy and palaeoecology. Review of Palaeobotany and Palynology, 179, 17-43.
Spore assemblages from the Lower Devonian Xujiachong Formation from Qujing, Yunnan, China. Palaeontology, 55(3), 583-611.
Pennsylvanian (mid/late Bolsovian-Asturian) permineralised plant assemblages of the Pennant Sandstone Formation of southern Britain: Systematics and palaeoecology. Review of Palaeobotany and Palynology, 173, 23-45.
A high-precision U-Pb age constraint on the Rhynie Chert Konservat-Lagerstatte: time scale and other implications. J GEOL SOC LONDON, 168(4), 863-872.
Evolutionary development of the plant and spore wall.. AoB Plants, 2011, plr027. View this article in WRRO
UV-B absorbing pigments in spores: biochemical responses to shade in a high-latitude birch forest and implications for sporopollenin-based proxies of past environmental change. POLAR RESEARCH, 30.
Dryland plant communities in the Pennsylvanian (mid- to late Bolsovian) Winterbourne Formation of Bristol, southern Britain: Further evidence for taphonomic megabias. Review of Palaeobotany and Palynology, 166(3-4), 268-285.
Earth's earliest non-marine eukaryotes. Nature.
The invasion of the land by plants: when and where?. New Phytol, 188(2), 306-309.
Spore assemblages from the Lower Devonian 'Lower Old Red Sandstone' deposits of Arran, Scotland. EARTH ENV SCI T R SO, 100, 391-397.
Palaeogeographic and palaeoclimatic considerations based on Ordovician to Lochkovian vegetation. Geological Society Special Publication, 339, 49-58.
Wall ultrastructure in three species of the dispersed spore Emphanisporites from the Early Devonian. Review of Palaeobotany and Palynology.
SPORE WALL ULTRASTRUCTURE IN THE EARLY LYCOPSID LECLERCQIA (PROTOLEPIDODENDRALES) FROM THE LOWER DEVONIAN OF NORTH AMERICA: EVIDENCE FOR A FUNDAMENTAL DIVISION IN THE LYCOPSIDS. AM J BOT, 96(10), 1849-1860.
Studies of spore/pollen wall ultrastructure in fossil and living plants: A review of the subject area and the contribution of Bernard Lugardon. REV PALAEOBOT PALYNO, 156(1-2), 2-6.
Morphology and wall ultrastructure of the megaspore Lagenicula (Triletes) mixta (Winslow 1962) comb. nov from the Carboniferous (Early Mississippian: mid Tournaisian) of Ohio, USA. REV PALAEOBOT PALYNO, 156(1-2), 51-61.
Studies of spore/pollen wall ultrastructure in fossil and living plants: A homage to Bernard Lugardon, a talented palynologist. REV PALAEOBOT PALYNO, 156(1-2), 1-1.
Origin and Radiation of the Earliest Vascular Land Plants. SCIENCE, 324(5925), 353-353.
ULTRASTRUCTURE OF ENIGMATIC PHYTOCLASTS (BANDED TUBES) FROM THE SILURIAN-LOWER DEVONIAN: EVIDENCE FOR AFFINITIES AND ROLE IN EARLY TERRESTRIAL ECOSYSTEMS. PALAIOS, 24(3-4), 167-180.
ULTRASTRUCTURE OF DISPERSED AND IN SITU SPECIMENS OF THE DEVONIAN SPORE RHABDOSPORITES LANGII: EVIDENCE FOR THE EVOLUTIONARY RELATIONSHIPS OF PROGYMNOSPERMS. PALAEONTOLOGY, 52, 139-167.
When scum ruled the Earth. Planet Earth(WINTER), 12-13.
Plant spore walls as a record of long-term changes in ultraviolet-B radiation. NAT GEOSCI, 1(9), 592-596.
Palaeophytogeographical and palaeoecological implications of a miospore assemblage of earliest Devonian (Lochkovian) age from Saudi Arabia. PALAEOGEOGR PALAEOCL, 250(1-4), 237-254.
Rapid determination of spore chemistry using thermochemolysis gas chromatography-mass spectrometry and micro-Fourier transform infrared spectroscopy.. Photochem Photobiol Sci, 6(6), 689-694.
Morphology and wall ultrastructure of the megaspore Lagenicula (Triletes) variabilis (Window, 1962) Arioli et al. (2004) from the Lower Carboniferous of Ohio, USA. REV PALAEOBOT PALYNO, 144(3-4), 231-248.
Early to Middle Devonian miospores from northern Saudi Arabia. Revue de Micropaleontologie, 50(1), 27-57.
Spores of the Rhynie chert plant Aglaophyton (Rhynia) major (Kidston and Lang) D.S Edwards, 1986. REV PALAEOBOT PALYNO, 142(3-4), 229-250.
Spore assemblages from the Lower Devonian 'Lower Old Red Sandstone' deposits of the Rhynie outlier, Scotland. T ROY SOC EDIN-EARTH, 97, 167-211.
The origin and early evolution of land plants: Palynological evidence. PALYNOLOGY, 30, 230-231.
Half a century of palynology at the University of Sheffield. Geological Society Special Publication, 241, 259-279.
New directions in Palaeozoic palynology. Review of Palaeobotany and Palynology, 130(1-4), 1-15.
Palaeoecology and palaeophytogeography of the rhynie chert plants: evidence from integrated analysis of in situ and dispersed spores.. Proc Biol Sci, 271(1542), 985-992.
Fragments of the earliest land plants.. Nature, 425(6955), 282-285. View this article in WRRO
Morphology and wall ultrastructure in Devonian spores with bifurcate-tipped processes. INT J PLANT SCI, 163(3), 451-474.
Lower Devonian spore assemblages from the Arbuthnott Group at Canterland Den in the Midland Valley of Scotland. REV PALAEOBOT PALYNO, 118(1-4), 157-180.
Spore assemblages from a Silurian sequence in borehole Hawiyah-151 from saudi Arabia. GeoArabia Special Publication, 1, 116-133.
Cryptospores and trilete spores from the Llandovery, Nuayyim-2 borehole, Saudi Arabia. GeoArabia Special Publication, 1, 92-115.
The microfossil record of early land plants.. Philos Trans R Soc Lond B Biol Sci, 355(1398), 717-731.
A new plant assemblage (microfossil and megafossil) from the Lower Old Red Sandstone of the Anglo-Welsh Basin: its implications for the palaeoecology of early terrestrial ecosystems.. Rev Palaeobot Palynol, 109(3-4), 161-196.
Tetrads in sporangia and spore masses from the Upper Silurian and Lower Devonian of the Welsh Borderland. Botanical Journal of the Linnean Society, 130(2), 111-156.
Tetrads in sporangia and spore masses from the upper Silurian and Lower Devonian of the Welsh Borderland. BOT J LINN SOC, 130(2), 111-156.
Sporangia containing Scylaspora from the lower Devonian of the Welsh Borderland. PALAEONTOLOGY, 42, 67-81.
Ordovician land plants: Evidence and interpretation. Acta Universitatis Carolinae, Geologica, 43(1-2), 275-277.
Palynology, biostratigraphy, and depositional environments around the Ludlow-Pridoli boundary at Woodbury Quarry, Herefordshire, England. Bollettino della Societa Paleontologica Italiana, 38(2-3), 397-404.
Ultrastructure of laevigate hilate spores in sporangia and spore masses from the Upper Silurian and Lower Devonian of the Welsh Borderland. PHILOS T ROY SOC B, 353(1378), 1983-2004.
The Cosheston Group (Lower Old Red Sandstone) in southwest Wales: age, correlation and palaeobotanical significance. GEOL MAG, 135(3), 397-412.
Permanent dyads in sporangia and spore masses from the Lower Devonian of the Welsh Borderland. Botanical Journal of the Linnean Society, 127(2), 117-147.
Curation of exceptionally preserved early land plant microfossils: problems and solutions. Curator, 39(3), 209-216.
Ultrastructure of Synorisporites downtonensis and Retusotriletes cf. coronadus in spore masses from the Přídoli of the Welsh Borderland. Palaeontology, 39(3), 783-800.
Sporomorph assemblages from the 'lower old red sandstone' of Lorne, Scotland. Special Papers in Palaeontology, 55, 41-101.
Cryptospores from the type area of the Caradoc Series in southern Britain. Special Papers in Palaeontology, 55, 103-136.
“Phytodebris” from Scottish Silurian and Lower Devonian continental deposits. Review of Palaeobotany and Palynology, 84(3-4), 255-279.
Palynology of the ‘Lower Old Red Sandstone’ at Glen Coe, Scotland. Geological Magazine, 131(4), 563-566.
A land plant microfossil assemblage of Mid Silurian age from the Stonehaven Group, Scotland. Journal of Micropalaeontology, 12(1), 47-66.
Terrestrial plant microfossils from Silurian inliers of the Midland Valley of Scotland. Palaeontology, 36(1), 155-193.
A Lower Devonian sporomorph assemblage from the Midland Valley of Scotland. Transactions of the Royal Society of Edinburgh: Earth Sciences, 84(2), 117-136.
Colonial palynomorphs from the Upper Ordovician of north-eastern Iran: ‘thalli’, coenobial Chlorophyceae (Hydrodictyaceae) or cyanobacteria?. Palynology, 1-11.
Pellicle ultrastructure demonstrates that Moyeria is a fossil euglenid. Palynology, 1-11.
Early Terrestrialization: Transition from Algal to Bryophyte Grade, Advances in Photosynthesis and Respiration (pp. 9-28). Springer Netherlands
Miospores and the emergence of land plants In Webby BD, Paris F, Droser ML & Percival IG (Ed.), The Great Ordovician Biodiversification Event (pp. 361-366).
Origin, function and development of the spore wall in early land plants, The Evolution of Plant Physiology (pp. 43-63). Elsevier
Origin, function and development of the spore wall in early land plants, The Evolution of Plant Physiology (pp. 43-63).
Embryophytes on land: The Ordovician to Lochkovian (Lower Devonian) record In Gensel PG & Edwards D (Ed.), Plants invade the land: Evolutionary and environmental perspectives (pp. 3-28).
Extracting plant mesofossils and megafossils by bulk acid maceration In Jones TP & Rowe NP (Ed.), Fossil plants and spores: modern techniques (pp. 11-14). Geological Society, London
The fossil record of early land plants and interrelationships between primitive embryophytes: too little and too late? In Bates JW, Ashton NW & Duckett JG (Ed.), Bryology for the Twenty-first Century (pp. 15-43). British Bryological Society
Older plant macerals (excluding spores) In Jansonius J & McGregor DC (Ed.), Palynology: principals and applications (pp. 383-387). AASP Foundation
How Advances in Phylogenetic Methods Change Our Understanding of the Evolutionary Emergence of Land Plants (Embryophyta) (pp. 1-11). Wiley
Conference proceedings papers