Professor Rob Freckleton
School of Biosciences
Chair in Population Biology, Head of School
  
  +44 114 222 0017
Full contact details
School of Biosciences
C214
Alfred Denny Building
Western Bank
Sheffield
S10 2TN
- Profile
 - 
    
- (2021-present) Head of School, School of Biosciences, University of Sheffield
 - (2018-present) Professor of Population Biology, School of Biosciences, University of Sheffield
 - (2012-2017) Director of Research and Innovation, Faculty of Science, University of Sheffield
 - (2006–2012) Royal Society University Research Fellow and Professor of Population Biology, Animal and Plant Sciences, University of Sheffield
 - (2004–2006) Royal Society University Research Fellow, University of Oxford
 - (2002–2004) Departmental Lecturer, University of Oxford
 - (2000–2002) Postdoctoral Researcher, University of Oxford
 - (1998–2000) Postdoctoral Researcher, University of East Anglia
 
 
- Research interests
 - 
    
My research focuses on modelling population and community dynamics. I am particularly interested in large scale population dynamics, although have a range of interests, including:
- Plant population ecology, modelling plant populations, modelling weed populations.
 - Evolutionary ecology, phylogenetic comparative methodology and its application to ecological problems.
 - Theoretical ecology, statistical methodology.
 
 
- Publications
 - 
    
Show: Featured publications All publications
Featured publications
Journal articles
-  Phylogenetic and Trait-Based Prediction of Extinction Risk for Data-Deficient Amphibians. Current Biology, 29, 1557-1563. View this article in WRRO 
					
					
				 -  Global pattern of nest predation is disrupted by climate change in shorebirds. Science, 362(6415), 680-683. View this article in WRRO 
					
					
				 -  The factors driving evolved herbicide resistance at a national scale. Nature Ecology and Evolution, 2, 529-536. View this article in WRRO 
					
					
				 -  Variation in helper effort among cooperatively breeding bird species is consistent with Hamilton's Rule.. Nature Communications, 7, 12663. View this article in WRRO 
					
					
				 -  C4 photosynthesis boosts growth by altering physiology, allocation and size.. Nature Plants, 2. View this article in WRRO 
					
					
				 -  The evolution of parental cooperation in birds. Proceedings of the National Academy of Sciences , 112(44), 13603-13608. View this article in WRRO 
					
					
				 -  Divorce and infidelity are associated with skewed adult sex ratios in birds.. Curr Biol, 24(8), 880-884. 
					
					
				 -  Pathogens and insect herbivores drive rainforest plant diversity and composition.. Nature, 506(7486), 85-88. 
					
					
				 -  The evolution of sex roles in birds is related to adult sex ratio.. Nat Commun, 4, 1587. 
					
					
				 -  Unraveling the life history of successful invaders.. Science, 337(6094), 580-583. 
					
					
				 -  Density-structured models for plant population dynamics.. Am Nat, 177(1), 1-17. 
					
					
				 -  Testing the Janzen-Connell mechanism: pathogens cause overcompensating density dependence in a tropical tree.. Ecol Lett, 13(10), 1262-1269. 
					
					
				 -  Niches versus neutrality: uncovering the drivers of diversity in a species-rich community.. Ecol Lett, 12(10), 1079-1090. 
					
					
				 -  Space versus phylogeny: disentangling phylogenetic and spatial signals in comparative data.. Proc Biol Sci, 276(1654), 21-30. 
					
					
				 -  Relating traits to diversification: a simple test.. Am Nat, 172(1), 102-115. 
					
					
				 
All publications
Journal articles
-  Tropical biodiversity loss from land-use change is severely underestimated by local-scale assessments. Nature Ecology & Evolution, 9(9), 1643-1655. View this article in WRRO 
					
					
				 -  Tropical land‐use change disrupts zeta‐diversity across taxa. Global Change Biology, 31(5). View this article in WRRO 
					
					
				 -  Global biogeography and macroevolutionary patterns of angiosperm inflorescence types. New Phytologist, 247(1), 358-368. View this article in WRRO 
					
					
				 -  Metabolic niches and plasticity of sand-dune plant communities along a trans-European gradient. Metabolites, 15(4). View this article in WRRO 
					
					
				 -  War‐induced behavioral change in spotted hyena (Crocuta crocuta) populations in Tigray, Northern Ethiopia. Ecology and Evolution, 15(2). View this article in WRRO 
					
					
				 -  Comparative analyses and phylogenetic dependence in traits and trends of the Dipterocarpaceae. Ecology and Evolution, 15(1). View this article in WRRO 
					
					
				 -  Black-grass monitoring using hyperspectral image data is limited by between-site variability. Remote Sensing, 16(24). View this article in WRRO 
					
					
				 -  Anisogamy and sex roles: a commentary. Evolution Letters, 8(6), 761-763. View this article in WRRO 
					
					
				 -  Pre‐copulatory sexual selection predicts sexual size dimorphism: a meta‐analysis of comparative studies. Ecology Letters, 27(9). View this article in WRRO 
					
					
				 -  Consequences of Local Conspecific Density Effects for Plant Diversity and Community Dynamics. Ecology Letters, 27(9). 
					
					
				 -  Species richness and evenness of European bird communities show differentiated responses to measures of productivity. Journal of Animal Ecology, 93(9), 1212-1224. View this article in WRRO 
					
					
				 -  The evolution of exceptional diversity in parental care and fertilization modes in ray-finned fishes. Evolution, 78(10), 1673-1684. View this article in WRRO 
					
					
				 -  Revealing uncertainty in the status of biodiversity change. Nature, 628, 788-794. View this article in WRRO 
					
					
				 -  Acting pre-emptively reduces the long-term costs of managing herbicide resistance. Scientific Reports, 14(1). View this article in WRRO 
					
					
				 -  Transformation of natural habitat disrupts biogeographical patterns of orchid diversity. Biological Conservation, 292. View this article in WRRO 
					
					
				 -  Tree diversity and liana infestation predict acoustic diversity in logged tropical forests. Biological Conservation, 291, 110488-110488. 
					
					
				 -  Quantifying the impacts of management and herbicide resistance on regional plant population dynamics in the face of missing data. Journal of Applied Ecology, 61(5), 1109-1126. View this article in WRRO 
					
					
				 -  Enhanced weathering in the US Corn Belt delivers carbon removal with agronomic benefits. Proceedings of the National Academy of Sciences, 121(9). View this article in WRRO 
					
					
				 -  Monitoring lianas from space: using Sentinel-2 imagery to observe liana removal in logged tropical forests. Forest Ecology and Management, 554. View this article in WRRO 
					
					
				 -  Evolution of reproductive modes in sharks and rays. Journal of Evolutionary Biology, 36(11), 1630-1640. View this article in WRRO 
					
					
				 -  Global distribution and evolutionary transitions of floral symmetry in angiosperms. Science Advances, 9(43). View this article in WRRO 
					
					
				 -  Evolutionary history and climate co‐determine the geographical variation in pollination modes of angiosperms in China. Global Ecology and Biogeography, 32(12), 2189-2198. 
					
					
				 -  Egg characteristics vary longitudinally in Arctic shorebirds. iScience, 26(6). View this article in WRRO 
					
					
				 -  Distance and density dependence in two native Bornean dipterocarp species. Ecology and Evolution, 13(4). View this article in WRRO 
					
					
				 -  Phylogeny and morphology determine vulnerability to global warming in Pristimantis frogs. Land, 12(1). 
					
					
				 -  High sensitivity of tropical forest birds to deforestation at lower altitudes. Ecology, 104(1). View this article in WRRO 
					
					
				 -  Crop diversification and parasitic weed abundance: a global meta-analysis. Scientific Reports, 12(1). 
					
					
				 -  The evolution of parental care in salamanders. Scientific Reports, 12(1). 
					
					
				 -  Quantification of CO2 removal in a large-scale enhanced weathering field trial on an oil palm plantation in Sabah, Malaysia. Frontiers in Climate, 4. View this article in WRRO 
					
					
				 -  A deep learning application to map weed spatial extent from unmanned aerial vehicles imagery. Remote Sensing, 14(17). 
					
					
				 -  Effects of density, species interactions, and environmental stochasticity on the dynamics of British bird communities. Ecology, 103(8). 
					
					
				 -  Dissecting weed adaptation: fitness and trait correlations in herbicide resistant Alopecurus myosuroides. Pest Management Science, 78(7), 3039-3050. View this article in WRRO 
					
					
				 -  Removing climbers more than doubles tree growth and biomass in degraded tropical forests. Ecology and Evolution, 12(3). View this article in WRRO 
					
					
				 -  Asking the right questions in explaining tropical diversity: response to Cannon and Lerdau. Trends in Ecology & Evolution, 37(5), 394-394. 
					
					
				 -  Drought exposure leads to rapid acquisition and inheritance of herbicide resistance in the weed Alopecurus myosuroides. Ecology and Evolution, 12(2). 
					
					
				 -  Exceptionally high apparent adult survival in three tropical species of plovers in Madagascar. Journal of Avian Biology, 2022(1). 
					
					
				 -  Animal migration to northern latitudes: environmental changes and increasing threats. Trends in Ecology & Evolution, 37(1), 30-41. 
					
					
				 -  Identifying existing management practices in the control of Striga asiatica within rice–maize systems in mid‐west Madagascar. Ecology and Evolution, 11(19), 13579-13592. 
					
					
				 -  Degree of anisogamy is unrelated to the intensity of sexual selection. Scientific Reports, 11(1). 
					
					
				 -  Global distribution and evolutionary transitions of angiosperm sexual systems. Ecology Letters, 24(9), 1835-1847. 
					
					
				 -  Evolution of large males is associated with female‐skewed adult sex ratios in amniotes. Evolution, 75(7), 1636-1649. 
					
					
				 -  Are evolutionary transitions in sexual size dimorphism related to sex determination in reptiles?. Journal of Evolutionary Biology, 34(4), 594-603. View this article in WRRO 
					
					
				 -  Developing hierarchical density‐structured models to study the national‐scale dynamics of an arable weed. Ecological Monographs, 91(3). View this article in WRRO 
					
					
				 -  Asking the Wrong Question in Explaining Tropical Diversity. Trends in Ecology & Evolution, 36(6), 482-484. 
					
					
				 -  Characterizing the environmental drivers of the abundance and distribution of Alopecurus myosuroides on a national scale. Pest Management Science, 77(6), 2726-2736. View this article in WRRO 
					
					
				 -  Overfishing and habitat loss drive range contraction of iconic marine fishes to near extinction. Science Advances, 7(7). 
					
					
				 -  Publisher Correction: Evolution of generalist resistance to herbicide mixtures reveals a trade-off in resistance management. Nature Communications, 11(1). 
					
					
				 -  Limited contributions of plant pathogens to density‐dependent seedling mortality of mast fruiting Bornean trees. Ecology and Evolution, 10(23), 13154-13164. View this article in WRRO 
					
					
				 -  Mapping the drivers of parasitic weed abundance at a national scale : a new approach applied to Striga asiatica in the mid‐west of Madagascar. Weed Research, 60(5), 323-333. View this article in WRRO 
					
					
				 -  Thermal tolerance and the importance of microhabitats for Andean frogs in the context of land use and climate change. Journal of Animal Ecology, 89(11), 2451-2460. 
					
					
				 -  C4 photosynthesis and the economic spectra of leaf and root traits independently influence growth rates in grasses. Journal of Ecology, 108(5), 1899-1909. View this article in WRRO 
					
					
				 -  Spatial patterns and drivers of angiosperm sexual systems in China differ between woody and herbaceous species. Frontiers in Plant Science, 11. 
					
					
				 -  Climate and mating systems as drivers of global diversity of parental care in frogs. Global Ecology and Biogeography, 29(8), 1373-1386. 
					
					
				 -  Evolution of generalist resistance to herbicide mixtures reveals a trade-off in resistance management. Nature Communications, 11(1). View this article in WRRO 
					
					
				 -  Ten years of Methods in Ecology and Evolution. Methods in Ecology and Evolution, 11(1), 4-5. 
					
					
				 -  Comparing life histories across taxonomic groups in multiple dimensions : how mammal-like are insects?. The American Naturalist, 195(1), 70-81. 
					
					
				 -  The costs of human-induced evolution in an agricultural system. Nature Sustainability, 3(1), 63-71. 
					
					
				 -  Drivers of large‐scale geographical variation in sexual systems of woody plants. Global Ecology and Biogeography, 29(3), 546-557. 
					
					
				 -  The circular nature of recurrent life cycle events: a test comparing tropical and temperate phenology. Journal of Ecology, 108(2), 393-404. 
					
					
				 -  European mushroom assemblages are darker in cold climates. Nature Communications, 10(1). 
					
					
				 -  Estimating the farm-level economic costs of spring cropping to manage Alopecurus myosuroides (black-grass) in UK agriculture. The Journal of Agricultural Science, 157(04), 318-332. 
					
					
				 -  Phylogeny and ecological processes influence grass coexistence at different spatial scales within the steppe biome. Oecologia, 191(1), 25-38. View this article in WRRO 
					
					
				 -  Comparative analysis of experimental data. Methods in Ecology and Evolution, 10(8), 1308-1321. View this article in WRRO 
					
					
				 -  Evolutionary epidemiology predicts the emergence of glyphosate resistance in a major agricultural weed. New Phytologist, 223(3), 1584-1594. View this article in WRRO 
					
					
				 -  Testing the ability of Unmanned Aerial Systems and machine learning to map weeds at subfield scales: a test with the weed Alopecurus myosuroides (Huds).. Pest Management Science, 75(8), 2283-2294. View this article in WRRO 
					
					
				 -  Response to Comment on “Global pattern of nest predation is disrupted by climate change in shorebirds”. Science, 364(6445). 
					
					
				 -  Phylogenetic and Trait-Based Prediction of Extinction Risk for Data-Deficient Amphibians. Current Biology, 29, 1557-1563. View this article in WRRO 
					
					
				 -  Testing mechanisms of compensatory fitness of dioecy in a cosexual world. Journal of Vegetation Science, 30(3), 413-426. 
					
					
				 -  Parental care and the evolution of terrestriality in frogs. Proceedings of the Royal Society B: Biological Sciences, 286(1900). View this article in WRRO 
					
					
				 -  Global pattern of nest predation is disrupted by climate change in shorebirds. Science, 362(6415), 680-683. View this article in WRRO 
					
					
				 -  Funder restrictions on application numbers lead to chaos. Trends in Ecology and Evolution, 33(8), 565-568. View this article in WRRO 
					
					
				 -  Defining and delivering resilient ecological networks: Nature conservation in England. Journal of Applied Ecology, 55(6), 2537-2543. 
					
					
				 -  The role of breeding system in community dynamics: growth and mortality in forests of different successional stages. Ecology and Evolution, 8(15), 7285-7296. View this article in WRRO 
					
					
				 -  Sex-biased breeding dispersal is predicted by social environment in birds. Ecology and Evolution, 8(13), 6483-6491. View this article in WRRO 
					
					
				 -  How to quantify competitive ability. Journal of Ecology, 106(5), 1902-1909. 
					
					
				 -  Abiotic and biotic determinants of coarse woody productivity in temperate mixed forests. Science of The Total Environment, 630, 422-431. 
					
					
				 -  The factors driving evolved herbicide resistance at a national scale. Nature Ecology and Evolution, 2, 529-536. View this article in WRRO 
					
					
				 -  Evaluating the potential of unmanned aerial systems for mapping weeds at field scales: a case study with Alopecurus myosuroides. Weed Research, 58(1), 35-45. View this article in WRRO 
					
					
				 -  Accessibility, reusability, reliability: Improving the standards for publishing code in  Methods in Ecology and Evolution. Methods in Ecology and Evolution, 9(1), 4-6. 
					
					
				 -  A critical review of integrated grass weed management in Ireland. Irish Journal of Agricultural and Food Research, 57(1), 15-28. View this article in WRRO 
					
					
				 -  Measuring the effectiveness of management interventions at regional scales by integrating ecological monitoring and modelling. Pest Management Science, 74(10), 2287-2295. View this article in WRRO 
					
					
				 -  Sex Allocation Patterns across Cooperatively Breeding Birds Do Not Support Predictions of the Repayment Hypothesis
. American Naturalist, 190(4), 547-556. View this article in WRRO 
					
					
				 -  Climate change mitigation : potential benefits and pitfalls of enhanced rock weathering in tropical agriculture.. Biology Letters, 13(4). View this article in WRRO 
					
					
				 -  Complex Relationships between Competing Guilds along Large-Scale Environmental Gradients. American Naturalist, 189(4), 407-421. View this article in WRRO 
					
					
				 -  Small-scale and regional spatial dynamics of an annual plant with contrasting sexual systems. Journal of Ecology, 105(4), 1044-1057. View this article in WRRO 
					
					
				 -  Trait Evolution in Adaptive Radiations: Modeling and Measuring Interspecific Competition on Phylogenies. The American Naturalist, 189(2), 121-137. View this article in WRRO 
					
					
				 -  Thermally buffered microhabitats recovery in tropical secondary forests following land abandonment. Biological Conservation, 201, 385-395. View this article in WRRO 
					
					
				 -  Variation in helper effort among cooperatively breeding bird species is consistent with Hamilton's Rule.. Nature Communications, 7, 12663. View this article in WRRO 
					
					
				 -  Special feature: 5th anniversary of Methods in Ecology and Evolution. Methods in Ecology and Evolution, 7(6), 634-635. 
					
					
				 -  A cautionary note on the use of Ornstein Uhlenbeck models in macroevolutionary studies. Biological Journal of the Linnean Society, 118(1), 64-77. View this article in WRRO 
					
					
				 -  C4 photosynthesis boosts growth by altering physiology, allocation and size.. Nature Plants, 2. View this article in WRRO 
					
					
				 -  Uncovering the spatio‐temporal drivers of species trait variances: a case study of Magnoliaceae in China. Journal of Biogeography, 43(6), 1179-1191. 
					
					
				 -  A Synthesis is Emerging between Biodiversity–Ecosystem Function and Ecological Resilience Research: Reply to Mori. Trends in Ecology & Evolution, 31(2), 89-92. 
					
					
				 -  The evolution of parental cooperation in birds. Proceedings of the National Academy of Sciences , 112(44), 13603-13608. View this article in WRRO 
					
					
				 -  Sex differences in parental care: Gametic investment, sexual selection, and social environment. Evolution, 69(11), 2862-2875. 
					
					
				 -  Biodiversity and Resilience of Ecosystem Functions. Trends in Ecology & Evolution, 30(11), 673-684. 
					
					
				 -  Weed Research – our aims and editorial policies. Weed Research, 55(5), 437-440. 
					
					
				 -  Phylogenetic constraints and trait correlates of flowering phenology in the angiosperm flora of 
C hina. Global Ecology and Biogeography, 24(8), 928-938.
					
				 -  Towards a general framework for predicting threat status of data-deficient species from phylogenetic, spatial and environmental information. Philosophical Transactions B: Biological Sciences, 370(1662). View this article in WRRO 
					
					
				 -  Agricultural Weed Research: A Critique and Two Proposals. Weed Science, 62(4), 672-678. 
					
					
				 -  Identifying the effect of density dependence, agricultural practices and climate variables on the long‐term dynamics of weed populations. Weed Research, 54(6), 556-564. 
					
					
				 -  Sex-biased survival predicts adult sex ratio variation in wild birds.. Proc Biol Sci, 281(1788), 20140342. 
					
					
				 -  Divorce and infidelity are associated with skewed adult sex ratios in birds.. Curr Biol, 24(8), 880-884. 
					
					
				 -  Links between plant species' spatial and temporal responses to a warming climate.. Proc Biol Sci, 281(1779), 20133017. 
					
					
				 -  Pathogens and insect herbivores drive rainforest plant diversity and composition.. Nature, 506(7486), 85-88. 
					
					
				 -  Testing the roles of competition, facilitation and stochasticity on community structure in a species-rich assemblage. Journal of Ecology. 
					
					
				 -  Mechanisms driving an unusual latitudinal diversity gradient for grasses. Global Ecology and Biogeography. 
					
					
				 -  Testing for enemy-mediated density-dependence in the mortality of seedlings: Field experiments with five Neotropical tree species. Oikos. 
					
					
				 -  Dynamic species distribution models from categorical survey data. Journal of Animal Ecology, 82(6), 1215-1226. 
					
					
				 -  What do we need to know to enhance the environmental sustainability of agricultural production? A prioritisation of knowledge needs for the UK food system. Sustainability Switzerland, 5(7), 3095-3115. 
					
					
				 -  Why care? Inferring the evolution of complex social behaviour.. J Evol Biol, 26(7), 1381-1391. 
					
					
				 -  Comment on “Bateman in Nature: Predation on Offspring Reduces the Potential for Sexual Selection”. Science, 340(6132), 549-549. 
					
					
				 -  Comment on "Bateman in nature: predation on offspring reduces the potential for sexual selection".. Science, 340(6132), 549. 
					
					
				 -  Empirical test of an agricultural landscape model: The importance of farmer preference for risk aversion and crop complexity. Sage Open, 3(2), 1-16. 
					
					
				 -  The evolution of sex roles in birds is related to adult sex ratio.. Nat Commun, 4, 1587. 
					
					
				 -  Interspecific differences in stochastic population dynamics explains variation in Taylor's temporal power law. Oikos. 
					
					
				 -  Identification of 100 fundamental ecological questions. Journal of Ecology, 101(1), 58-67. 
					
					
				 -  Spatial and temporal variability in positive and negative plant-bryophyte interactions along a latitudinal gradient. Journal of Ecology, 101(2), 465-474. 
					
					
				 -  Unraveling the life history of successful invaders.. Science, 337(6094), 580-583. 
					
					
				 -  Fast likelihood calculations for comparative analyses. METHODS IN ECOLOGY AND EVOLUTION, 3(5), 940-947. 
					
					
				 -  Consequences of changing rainfall for fungal pathogen-induced mortality in tropical tree seedlings.. Ecol Evol, 2(7), 1408-1413. 
					
					
				 -  Phylogenetic niche conservatism in C4 grasses.. Oecologia, 170(3), 835-845. 
					
					
				 -  Megacycles of atmospheric carbon dioxide concentration correlate with fossil plant genome size.. Philos Trans R Soc Lond B Biol Sci, 367(1588), 556-564. 
					
					
				 -  Assessing the role of competition and stress: A critique of importance indices and the development of a new approach. Journal of Ecology, 100(3), 577-585. 
					
					
				 -  An objective, niche-based approach to indicator species selection. METHODS IN ECOLOGY AND EVOLUTION, 3(2), 317-326. 
					
					
				 -  Making predictive ecology more relevant to policy makers and practitioners.. Philos Trans R Soc Lond B Biol Sci, 367(1586), 322-330. 
					
					
				 -  Environmental factors determining the phylogenetic structure of C4 grass communities. JOURNAL OF BIOGEOGRAPHY, 39(2), 232-246. 
					
					
				 -  Changes in the large-scale distribution of plants: Extinction, colonisation and the effects of climate. Journal of Ecology, 100(2), 519-529. 
					
					
				 -  MOTMOT: models of trait macroevolution on trees. METHODS IN ECOLOGY AND EVOLUTION, 3(1), 145-151. 
					
					
				 -  Characterizing abundance-occupancy relationships: There is no artefact. Global Ecology and Biogeography. 
					
					
				 -  Phylogenetic conservatism of environmental niches in mammals.. Proc Biol Sci, 278(1716), 2384-2391. 
					
					
				 -  Problems formalising the concept of importance in ecology.. Trends Ecol Evol, 26(10), 498-499. 
					
					
				 -  Comparative methods as a statistical fix: the dangers of ignoring an evolutionary model.. Am Nat, 178(1), E10-E17. 
					
					
				 -  GrassPortal: An online ecological and evolutionary data facility for the grasses. S AFR J BOT, 77(2), 566-566. 
					
					
				 -  Understanding geographical patterns of grass species richness - The global and local context. S AFR J BOT, 77(2), 565-566. 
					
					
				 -  Problems formalising the concept of importance in ecology. Trends in Ecology and Evolution. 
					
					
				 -  Density-structured models for plant population dynamics.. Am Nat, 177(1), 1-17. 
					
					
				 -  From meso- to macroscale population dynamics: a new density-structured approach. METHODS ECOL EVOL, 2(3), 289-302. 
					
					
				 -  Dealing with collinearity in behavioural and ecological data: Model averaging and the problems of measurement error. Behavioral Ecology and Sociobiology, 65(1), 91-101. 
					
					
				 -  Rarity, life history and scaling of the dynamics in time and space of British birds.. J Anim Ecol, 80(1), 215-224. 
					
					
				 -  Model averaging, missing data and multiple imputation: A case study for behavioural ecology. Behavioral Ecology and Sociobiology, 65(1), 103-116. 
					
					
				 -  Constant final yield. Annual Review of Ecology Evolution and Systematics, 41, 173-192. 
					
					
				 -  Phylogenetic comparative approaches for studying niche conservatism.. J Evol Biol, 23(12), 2529-2539. 
					
					
				 -  Testing the Janzen-Connell mechanism: pathogens cause overcompensating density dependence in a tropical tree.. Ecol Lett, 13(10), 1262-1269. 
					
					
				 -  Who cares? Quantifying the evolution of division of parental effort. METHODS ECOL EVOL, 1(3), 221-230. 
					
					
				 -  Can phylogenetics identify C(4) origins and reversals?. Trends Ecol Evol, 25(7), 403-409. 
					
					
				 -  The origins of C4 grasslands: integrating evolutionary and ecosystem science.. Science, 328(5978), 587-591. 
					
					
				 -  Understanding the role of species dynamics in abundance-occupancy relationships. J ECOL, 98(3), 645-658. 
					
					
				 -  Methods in Ecology and Evolution. METHODS ECOL EVOL, 1(1), 1-2. 
					
					
				 -  Model averaging, missing data and multiple imputation: a case study for behavioural ecology. Behavioral Ecology and Sociobiology, 1-14. 
					
					
				 -  Dealing with collinearity in behavioural and ecological data: model averaging and the problems of measurement error. Behavioral Ecology and Sociobiology, 1-11. 
					
					
				 -  Niches versus neutrality: uncovering the drivers of diversity in a species-rich community.. Ecol Lett, 12(10), 1079-1090. 
					
					
				 -  Critical parameters for predicting population fluctuations of some British passerines.. J Anim Ecol, 78(5), 1063-1075. 
					
					
				 -  The seven deadly sins of comparative analysis.. J Evol Biol, 22(7), 1367-1375. 
					
					
				 -  Ecological selection pressures for C4 photosynthesis in the grasses.. Proc Biol Sci, 276(1663), 1753-1760. 
					
					
				 -  Predictive models of weed population dynamics. WEED RES, 49(3), 225-232. 
					
					
				 -  Measuring the importance of competition in plant communities. J ECOL, 97(3), 379-384. 
					
					
				 -  Seed mass, abundance and breeding system among tropical forest species: do dioecious species exhibit compensatory reproduction or abundances?. J ECOL, 97(3), 555-566. 
					
					
				 -  Habitat associations of British breeding farmland birds. BIRD STUDY, 56, 43-52. 
					
					
				 -  Are parental care trade-offs in shorebirds driven by parental investment or sexual selection?. J Evol Biol, 22(4), 672-682. 
					
					
				 -  Integrating socio-economics and ecology: a taxonomy of quantitative methods and a review of their use in agro-ecology. J APPL ECOL, 46(2), 269-277. 
					
					
				 -  Space versus phylogeny: disentangling phylogenetic and spatial signals in comparative data.. Proc Biol Sci, 276(1654), 21-30. 
					
					
				 -  Missing inaction: the dangers of ignoring missing data.. Trends Ecol Evol, 23(11), 592-596. 
					
					
				 -  Environment, migratory tendency, phylogeny and basal metabolic rate in birds.. PLoS One, 3(9), e3261. 
					
					
				 -  Modelling the effects of management on population dynamics: some lessons from annual weeds. J APPL ECOL, 45(4), 1050-1058. 
					
					
				 -  Relating traits to diversification: a simple test.. Am Nat, 172(1), 102-115. 
					
					
				 -  Does double-blind review benefit female authors?. Trends Ecol Evol, 23(7), 351-353. 
					
					
				 -  Future novel threats and opportunities facing UK biodiversity identified by horizon scanning. J APPL ECOL, 45(3), 821-833. 
					
					
				 -  What use is an infertile sperm? A comparative study of sperm-heteromorphic Drosophila.. Evolution, 62(2), 374-385. 
					
					
				 -  Parental conflict in birds: comparative analyses of offspring development, ecology and mating opportunities.. Proc Biol Sci, 275(1632), 301-307. 
					
					
				 -  Only half right: species with female-biased sexual size dimorphism consistently break Rensch's rule.. PLoS One, 2(9), e897. 
					
					
				 -  The role of ecological theory in microbial ecology.. Nat Rev Microbiol, 5(5), 384-392. 
					
					
				 -  Introducing Population Ecology: Key Concepts And Current Research Directions. Ecology, 88(4), 1070-1072. 
					
					
				 -  Biogeographical basis of recent phenotypic divergence among birds: a global study of subspecies richness.. Evolution, 61(4), 942-957. 
					
					
				 -  Phylogenetic evidence for deleterious mutation load in RNA viruses and its contribution to viral evolution.. Mol Biol Evol, 24(3), 845-852. 
					
					
				 -  Should conservation strategies consider spatial generality? Farmland birds show regional not national patterns of habitat association.. Ecol Lett, 10(1), 25-35. 
					
					
				 -  Abundance-occupancy dynamics in a human dominated environment: linking interspecific and intraspecific trends in British farmland and woodland birds.. J Anim Ecol, 76(1), 123-134. 
					
					
				 -  Detecting non-Brownian trait evolution in adaptive radiations.. PLoS Biol, 4(11), e373. 
					
					
				 -  Pathogens, density dependence and the coexistence of tropical trees.. Proc Biol Sci, 273(1604), 2909-2916. 
					
					
				 -  Why do we still use stepwise modelling in ecology and behaviour?. J Anim Ecol, 75(5), 1182-1189. 
					
					
				 -  The identification of 100 ecological questions of high policy relevance in the UK. J APPL ECOL, 43(4), 617-627. 
					
					
				 -  Census error and the detection of density dependence.. J Anim Ecol, 75(4), 837-851. 
					
					
				 -  Ecology predicts large-scale patterns of phylogenetic diversification in birds.. Am Nat, 168(2), 220-229. 
					
					
				 -  Special profile: Making better biogeographical predictions of species' distributions. Journal of Applied Ecology, 43(3), 385. 
					
					
				 -  Comparative analyses of the influence of developmental mode on phenotypic diversification rates in shorebirds.. Proc Biol Sci, 273(1594), 1619-1624. 
					
					
				 -  Plant pathogens drive density-dependent seedling mortality in a tropical tree.. Ecol Lett, 9(5), 569-574. 
					
					
				 -  Phenotypic plasticity in the scaling of avian basal metabolic rate.. Proc Biol Sci, 273(1589), 931-937. 
					
					
				 -  Erratum: Large-scale population dynamics, abundance-occupancy relationships and the scaling from local to regional population size (Journal of Animal Ecology (2005) 74, (353-364) DOI: 10.1111/j.1365-2656.2005.00931.x). Journal of Animal Ecology, 75(1), 314. 
					
					
				 -  UNUSUAL SPERM MORPHOLOGY IN THE EURASIAN BULLFINCH (PYRRHULA PYRRHULA). The Auk, 123(2), 383-383. 
					
					
				 -  Large-scale population dynamics, abundance-occupancy relationships and the scaling from local to regional population size (vol 74, pg 353, 2005). J ANIM ECOL, 75(1), 314-314. 
					
					
				 -  Distributions of habitat suitability and the abundance-occupancy relationship.. Am Nat, 167(2), 260-275. 
					
					
				 -  Ecology. Population dynamics: growing to extremes.. Science, 309(5734), 567-568. 
					
					
				 -  Habitat selection by yellowhammers Emberiza citrinella on lowland farmland at two spatial scales: implications for conservation management. J APPL ECOL, 42(2), 270-280. 
					
					
				 -  Large-scale population dynamics, abundance-occupancy relationships and the scaling from local to regional population size. J ANIM ECOL, 74(2), 353-364. 
					
					
				 -  The changing face of applied ecology. J APPL ECOL, 42(1), 1-3. 
					
					
				 -  Laws of Population Ecology. Ecology, 86(1), 271-272. 
					
					
				 -  Sexual selection explains Rensch's rule of size dimorphism in shorebirds.. Proc Natl Acad Sci U S A, 101(33), 12224-12227. 
					
					
				 -  The problems of prediction and scale in applied ecology: the example of fire as a management tool. J APPL ECOL, 41(4), 599-603. 
					
					
				 -  Seeing the jungle for the trees. Journal of Biogeography, 31(8), 1377-1377. 
					
					
				 -  The Future of GM Crops. Science, 304(5673), 959-963. 
					
					
				 -  The Future of GM Crops [2] (multiple letters). Science, 304(5673). 
					
					
				 -  The future of GM crops - Response. SCIENCE, 304(5673), 960-961. 
					
					
				 -  Coral reef cascades and the indirect effects of predator removal by exploitation. ECOL LETT, 7(5), 410-416. 
					
					
				 -  Erratum: Demographic threats to the sustainability of Brazil nut exploitation (Science (December 19) (2112)). Science, 303(5661), 1138. 
					
					
				 -  Amelioration of biodiversity impacts of genetically modified crops: predicting transient versus long-term effects.. Proc Biol Sci, 271(1536), 325-331. 
					
					
				 -  Demographic threats to the sustainability of Brazil nut exploitation.. Science, 302(5653), 2112-2114. 
					
					
				 -  Predicting the response of farmland bird populations to changing food supplies. J APPL ECOL, 40(6), 970-983. 
					
					
				 -  Ecology. Deciding the future of GM crops in Europe.. Science, 302(5647), 994-996. 
					
					
				 -  Predicting the impacts of harvesting using structured population models: the importance of density-dependence and timing of harvest for a tropical palm tree. J APPL ECOL, 40(5), 846-858. 
					
					
				 -  Bergmann's rule and body size in mammals.. Am Nat, 161(5), 821-825. 
					
					
				 -  Are all plant populations metapopulations?. J ECOL, 91(2), 321-324. 
					
					
				 -  Relative testis size and sperm morphometry across mammals: no evidence for an association between sperm competition and sperm length.. Proc Biol Sci, 270(1515), 625-632. 
					
					
				 -  Honesty and cheating in cleaning symbioses: evolutionarily stable strategies defined by variable pay-offs.. Proc Biol Sci, 270(1512), 299-305. 
					
					
				 -  Declines in the numbers of amateur and professional taxonomists: implications for conservation. ANIM CONSERV, 5, 245-249. 
					
					
				 -  Phylogenetic analysis and comparative data: a test and review of evidence.. Am Nat, 160(6), 712-726. 
					
					
				 -  Chaotic mating systems. TRENDS ECOL EVOL, 17(11), 493-495. 
					
					
				 -  Are weed population dynamics chaotic?. J APPL ECOL, 39(5), 699-707. 
					
					
				 -  Large-scale spatial dynamics of plants: metapopulations, regional ensembles and patchy populations. J ECOL, 90(3), 419-434. 
					
					
				 -  On the misuse of residuals in ecology: regression of residuals vs. multiple regression. J ANIM ECOL, 71(3), 542-545. 
					
					
				 -  Evolutionary transitions in parental care and live bearing in vertebrates.. Philos Trans R Soc Lond B Biol Sci, 357(1419), 269-281. 
					
					
				 -  Do in-hospital waiting lists show self-regulation?. J R Soc Med, 95(3), 164. 
					
					
				 -  Predicting competition coefficients for plant mixtures: reciprocity, transitivity and correlations with life-history traits. ECOL LETT, 4(4), 348-357. 
					
					
				 -  Asymmetric competition between plant species. FUNCT ECOL, 15(5), 615-623. 
					
					
				 -  Hospital waiting-lists. Do power laws imply self-regulation?. Nature, 413(6854), 382. 
					
					
				 -  Nonmanipulative determination of plant community dynamics.. Trends Ecol Evol, 16(6), 301-307. 
					
					
				 -  On detecting and measuring competition in spatially structured plant communities. ECOL LETT, 3(5), 423-432. 
					
					
				 -  Phylogenetic tests of ecological and evolutionary hypotheses: checking for phylogenetic independence. FUNCT ECOL, 14(1), 129-134. 
					
					
				 -  Population dynamics of Vulpia ciliata: regional, patch and local dynamics. J ECOL, 88(6), 1012-1029. 
					
					
				 -  Designs for greenhouse studies of interactions between plants: an analytical perspective. J ECOL, 88(3), 386-391. 
					
					
				 -  Predictions of biodiversity response to genetically modified herbicide-tolerant crops.. Science, 289(5484), 1554-1557. 
					
					
				 -  Determinants of the abundance of invasive annual weeds: community structure and non-equilibrium dynamics.. Proc Biol Sci, 267(1448), 1153-1161. 
					
					
				 -  Biological control as a learning process.. Trends Ecol Evol, 15(7), 263-264. 
					
					
				 -  The mis-measurement of plant competition. FUNCT ECOL, 13(2), 285-287. 
					
					
				 -  The population dynamics of Anisantha sterilis in winter wheat: comparative demography and the role of management. J APPL ECOL, 36(4), 455-471. 
					
					
				 -  The role of density dependence in the population dynamics of a tropical palm. ECOLOGY, 80(8), 2635-2650. 
					
					
				 -  A Course in Demographic Analysis. Ecology, 80(5), 1784-1785. 
					
					
				 -  A Course in Demographic Analysis. Ecology, 80(5), 1784-1784. 
					
					
				 -  Yield of sugar beet in relation to weather and nutrients. AGR FOREST METEOROL, 93(1), 39-51. 
					
					
				 -  What is the Allee effect?. OIKOS, 87(1), 185-190. 
					
					
				 -  Predicting the determinants of weed abundance: a model for the population dynamics of Chenopodium album in sugar beet. J APPL ECOL, 35(6), 904-920. 
					
					
				 -  How does temporal variability affect predictions of weed population numbers?. J APPL ECOL, 35(2), 340-344. 
					
					
				 -  Measuring plant neighbour effects. FUNCT ECOL, 11(4), 532-534. 
					
					
				 -  Measuring and modelling plant neighbour effects - Response. FUNCT ECOL, 11(4), 536-536. 
					
					
				 -  Measuring and modelling plant neighbour effects - Reply. FUNCTIONAL ECOLOGY, 11(4), 534-535. 
					
					
				 -  Measuring and modelling plant neighbour effects: A reply to Freckleton and Watkinson. Functional Ecology, 11(4), 534-536. 
					
					
				 -  Quantifying the impact of arbuscular mycorrhiza on plant competition. J ECOL, 85(4), 541-545. 
					
					
				 -  Applied Population Ecology: A Supply-Demand Approach.. The Journal of Applied Ecology, 33(6), 1588-1588. 
					
					
				 -  Author Correction: Degree of anisogamy is unrelated to the intensity of sexual selection. Scientific Reports, 11(1). 
					
					
				 -  The occurrence of herbicide-resistant Avena fatua (wild oats) populations to ACCase-inhibiting herbicides in Ireland. Irish Journal of Agricultural and Food Research, 60(1). 
					
					
				 -  The Epidemiological Framework for Biological Invasions (EFBI): an interdisciplinary foundation for the assessment of biosecurity threats. NeoBiota, 62, 161-192. 
					
					
				 -  commentary: A Darwinian approach to community ecology. Frontiers of Biogeography, 1(2). 
					
					
				 -  Ecophysiological traits of grasses: resolving the effects of photosynthetic pathway and phylogeny. Nature Precedings. 
					
					
				 
Book chapters
-  12. Transforming Practice, Transforming Conservation (pp. 367-386). Open Book Publishers 
					
					
				 -  2. Gathering and Assessing Pieces of Evidence, Transforming Conservation (pp. 31-74). Open Book Publishers 
					
					
				 -  12. Transforming Practice Checklists for Delivering Change, TRANSFORMING CONSERVATION (pp. 367-386). 
					
					
				 -  Conservation decisions in the face of uncertainty, Conservation Research, Policy and Practice (pp. 183-195). Cambridge University Press 
					
					
				 -  What Do We Need to Know to Enhance the Environmental Sustainability of Agricultural Production? A Prioritisation of Knowledge Needs for the UK Food System, Sustainable Agriculture and Food Supply Scientific Economic and Policy Enhancements (pp. 293-314). 
					
					
				 -  Recent advances in comparative methods, Social Behaviour (pp. 110-126). Cambridge University Press 
					
					
				 -  Stochasticty, nonlinearity and instability in biological invasions, Conceptual Ecology and Invasion Biology Reciprocal Approaches to Nature (pp. 125-146). 
					
					
				 -  Weed invasions of Australian farming systems: from ecology to economics, The Economics of Biological Invasions Edward Elgar Publishing 
					
					
				 
Conference proceedings
Preprints
-  War-induced behavioral change in spotted hyena (Crocuta crocuta) populations in Tigray, northern Ethiopia, Wiley. 
					
					
				 -  Enhanced weathering in the U.S. Corn Belt delivers carbon removal with agronomic benefits, arXiv. 
					
					
				 -  Managing herbicide resistance doesn't pay - but acting pre-emptively does., CABI Publishing. 
					
					
				 -  Overconfidence undermines global wildlife abundance trends, Cold Spring Harbor Laboratory. 
					
					
				 -  Identifying cultural, climatic and temporal factors influencing Striga asiatica abundance within rice--maize systems in mid-west Madagascar, Wiley. 
					
					
				 -  Trait evolution in adaptive radiations: modelling and measuring interspecific competition on phylogenies, Cold Spring Harbor Laboratory. 
					
					
				 -  Bias and measurement error in comparative analyses: a case study with the Ornstein Uhlenbeck model, Cold Spring Harbor Laboratory. 
					
					
				 -  Egg Characteristics Vary Longitudinally in Arctic Shorebirds. 
					
					
				 
 -  Phylogenetic and Trait-Based Prediction of Extinction Risk for Data-Deficient Amphibians. Current Biology, 29, 1557-1563. View this article in WRRO 
					
 
- Research group
 - 
    
Postdoctoral reseachers
- Dr Helen Hicks (2014-2017) Herbicide resistance in Blackgrass
 - Dr Daniel Bearup (2016-2017) Modelling soil quality in arable fields and landscapes
 - Dr Shaun Coutts (2015-2018) Modelling Blackgrass populations
 
PhD students
- Ronan Byrne (2016-2019) The spread of arable weeds in Ireland
 - Robert Goodsell (2014-2018) Monitoring farmland arthropod communities
 - James Lambert (2014-2018) Using UAVs to monitor weeds
 - Nazarin Mohammed (2015-2019) Modelling forest restoration
 - Pamela Gonzalez del Pliego Castaneda (2013-2017) Tropical conservation
 - Simon Rolph
 
Former group members
- Kwadjo Ahodo
 - Magnus Clark
 - Vian Hamed
 - Monica Ortiz
 - Dr Tom Webb Royal Society University Research Fellow at APS Sheffield
 - Dr Simon Queenborough Research fellow at NCEAS
 - Dr Colin Harrower Centre for Ecology and Hydrology
 - Dr Rémi Vergnon University of Waageningen
 
Collaborations
I also have collaborative projects with the following:
- Walter Jetz – phylogenetic & spatial analysis of comparative data (Yale: http://www.yale.edu/jetz/)
 - Tamas Szekely – Sexual conflict in birds (University of Bath http://www.bath.ac.uk/bio-sci/tamas.html)
 - John Pannell – Large scale dynamics and sexual systems in annual mercury (University of Oxford, Plant Sciences http://dps.plants.ox.ac.uk/external/)
 - Owen Lewis – Density-dependence and pathogens in tropical forests (University of Oxford, Zoology http://www.zoo.ox.ac.uk/ecology/owen_lewis.html)
 - Bill Sutherland – Developing models for arable farming systems (University of Cambridge).
 - Andrew Watkinson – Plant population and community ecology (University of East Anglia).
 
 
- Teaching activities
 - 
    
At Level 1 I lecture for APS121 (Evolution) and at Level 3 I lecture for APS342 (Evolutionary Ecology).
At level 3 I teach on a 10-day residential course in Borneo (APS 306 - Tropical Biology Rain Forest Field Course) and I supervise Projects (APS330) on herbivore-plant interactions, spatial ecology and plant population dynamics, usually field-based.
I have supervised Level 4 Research Projects (APS 406) on conservation, macroecology and evolution.
 
- Professional activities and memberships
 - 
    
- Academy of Finland Panel Chair (2017)
 - NERC Panel C member (2016)
 - Global Food Security Programme Panel member (2016)
 - BBSRC/Newton Joint Research Centres, Panel co-chair (2015)
 - NERC Neonicotinoid Experiment Project Advisory Group (2015-present)
 - DEFRA Rural Development Plan Expert Evaluation Panel (2014-present)
 - Founding Editor in Chief, Methods in Ecology & Evolution (2009-present)
 - FORMAS (Swedish Research Council) Grant Panel (2014-2017)
 - NERC Panel D Core Member (2013–present)
 - Senior Editor, Journal of Applied Ecology (2004–2009)
 - Academy of Finland Grant Review Panel (2009-present)
 - Swedish Academy Biodiversity Review (2010)
 - NERC Ecology and Hydrology Funding Initiative moderating panel (2008)
 - Professorial appointment panel, NTNU, Norway (2008)
 - NERC peer review college (2004-2007; 2011-2012).
 - NERC small and standard grants panels committee member (2005, 2012)
 - PhD external examiner (Imperial College, London; UCL; University of Birmingham; University of Aberdeen; University of Reading; University of Newcastle, Swedish Agricultural University; University of Lausanne)