Professor Ben J Hatchwell

School of Biosciences

Professor of Evolutionary Ecology

b.hatchwell@sheffield.ac.uk
+44 114 222 4625

B91, Alfred Denny Building

Full contact details

Professor Ben J Hatchwell
School of Biosciences
B91
Alfred Denny Building
Western Bank
Sheffield
S10 2TN

Profile

Professor of Evolutionary Ecology, University of Sheffield (2007-present)
Reader, University of Sheffield (2004 - 2006)
Senior Lecturer in Zoology, University of Sheffield (2002-2004)
Lecturer in Zoology, University of Sheffield (1993-2002)
Post-doc, University of Oxford (1991-1993)
Post-doc, University of Cambridge (1988-1991)
PhD (1988), University of Sheffield
BA (1984), University of Oxford

Research interests

Social Evolution

My principal research interest is in social evolution and reproductive strategies in birds. The main approach of my research is to use field observations and experiments to test evolutionary theory. I have been studying cooperative breeding in long-tailed tits since 1994, and have also collaborated recently on other projects on cooperative behaviour in sociable weavers, riflemen and monk parakeets. Specific research interests include:

The ecological and demographic factors that promote the evolution of cooperation in animal societies.
The influence of individual dispersal decisions on the genetic structure of populations and the consequences for cooperative behaviour
The fitness consequences of alternative reproductive strategies of individuals in cooperative groups
Mechanisms of kin recognition in social animals
Proximate and ultimate causes of variation in parental investment

Population Ecology

I am also interested in pure and applied aspects of avian population ecology, including long-term studies of seabirds, parrot conservation and the ecology of urban bird populations. Recent projects include studies of guillemots, blackbirds and three species of parrot. Specific areas of interest include:

The population dynamics of guillemots on Skomer Island
Population biology of vulnerable parrot populations
The relationship between urban and rural bird populations
Invasion biology of monk parakeets in Europe

Publications

Show: Featured publications All publications

Featured publications

Journal articles

Ellison N, Hatchwell B, Biddiscombe S, Napper C & Potts J (2020) Mechanistic home range analysis reveals drivers of space use patterns for a non-territorial passerine. Journal of Animal Ecology, 89(12), 2763-2776. View this article in WRRO
Leedale AE, Simeoni M, Sharp SP, Green JP, Slate J, Lachlan RF, Robinson EJH & Hatchwell BJ (2020) Cost, risk, and avoidance of inbreeding in a cooperatively breeding bird. Proceedings of the National Academy of Sciences, 117(27), 15724-15730.
Leedale AE, Lachlan RF, Robinson EJH & Hatchwell BJ (2020) Helping decisions and kin recognition in long-tailed tits : is call similarity used to direct help towards kin?. Philosophical Transactions of the Royal Society B: Biological Sciences, 375(1802). View this article in WRRO
Green J & Hatchwell BJ (2018) Inclusive fitness consequences of dispersal decisions in a cooperatively breeding bird, the long-tailed tit (Aegithalos caudatus). Proceedings of the National Academy of Sciences, 115(47), 12011-12016. View this article in WRRO
Leedale A, Sharp SP, Simeoni M, Robinson EJH & Hatchwell BJ (2018) Fine‐scale genetic structure and helping decisions in a cooperatively breeding bird . Molecular Ecology, 27(7), 1714-1726. View this article in WRRO
Khwaja N, Hatchwell BJ, Freckleton RP & Green JP (2017) Sex Allocation Patterns across Cooperatively Breeding Birds Do Not Support Predictions of the Repayment Hypothesis . American Naturalist, 190(4), 547-556. View this article in WRRO
Bebbington K & Hatchwell BJ (2016) Coordinated parental provisioning is related to feeding rate and reproductive success in a songbird. Behavioral Ecology, 27(2), 652-659.
Green JP, Freckleton RP & Hatchwell BJ (2016) Variation in helper effort among cooperatively breeding bird species is consistent with Hamilton’s Rule. Nature Communications, 7(-), ---.
Sæther B-E, Grøtan V, Engen S, Coulson T, Grant PR, Visser ME, Brommer JE, Rosemary Grant B, Gustafsson L, Hatchwell BJ , Jerstad K et al (2016) Demographic routes to variability and regulation in bird populations. Nature Communications, 7(-), ---.
Adams MJ, Robinson MR, Mannarelli M-E & Hatchwell BJ (2015) Social genetic and social environment effects on parental and helper care in a cooperatively breeding bird. Proceedings of the Royal Society B: Biological Sciences, 282(1810).
van Dijk RE, Kaden JC, Arguelles Tico A, Dawson DA, Burke T & Hatchwell BJ (2014) Cooperative investment in public goods is kin directed in communal nests of social birds. Ecology Letters, 17(9), 1141-1148.
Hatchwell BJ, Gullett PR & Adams MJ (2014) Helping in cooperatively breeding long-tailed tits: a test of Hamilton's rule.. Philos Trans R Soc Lond B Biol Sci, 369(1642), 20130565.
Abbot P, Abe J, Alcock J, Alizon S, Alpedrinha JA, Andersson M, Andre JB, van Baalen M, Balloux F, Balshine S , Barton N et al (2011) Inclusive fitness theory and eusociality.. Nature, 471(7339), E1-E4.
Meade J, Nam K-B, Beckerman AP & Hatchwell BJ (2010) Consequences of 'load-lightening' for future indirect fitness gains by helpers in a cooperatively breeding bird.. J Anim Ecol, 79(3), 529-537.
Hatchwell BJ (2010) Cryptic Kin Selection: Kin Structure in Vertebrate Populations and Opportunities for Kin-Directed Cooperation. ETHOLOGY, 116(3), 203-216.
Hatchwell BJ (2009) The evolution of cooperative breeding in birds: kinship, dispersal and life history.. Philos Trans R Soc Lond B Biol Sci, 364(1533), 3217-3227.
Evans KL, Gaston KJ, Frantz AC, Simeoni M, Sharp SP, McGowan A, Dawson DA, Walasz K, Partecke J, Burke T & Hatchwell BJ (2009) Independent colonization of multiple urban centres by a formerly forest specialist bird species.. Proc Biol Sci, 276(1666), 2403-2410.
Sharp SP, McGowan A, Wood MJ & Hatchwell BJ (2005) Learned kin recognition cues in a social bird.. Nature, 434(7037), 1127-1130.
MacColl ADC & Hatchwell BJ (2003) Heritability of parental effort in a passerine bird.. Evolution, 57(9), 2191-2195.
MacColl ADC & Hatchwell BJ (2003) HERITABILITY OF PARENTAL EFFORT IN A PASSERINE BIRD. Evolution, 57(9), 2191-2191.

All publications

Journal articles

Dawson Pell FSE, Senar JC, Ortega‐Segalerva A, Burke T & Hatchwell BJ (2025) Variable social organization and breeding system of a social parrot revealed by genetic analysis. Ibis. View this article in WRRO
Morinay J, Woodward BK, Russell AF, Sharp SP & Hatchwell BJ (2025) Correction to ‘Ecological and demographic drivers of kin‐directed cooperation in a social bird: Insights from a long‐term study’. Journal of Animal Ecology, 94(6), 1317-1317.
Baxter J, Baxter A & Hatchwell BJ (2025) Drivers of variation in egg size in a cooperative breeder with a redirected helping system. Journal of Evolutionary Biology, 38(8), 1041-1049. View this article in WRRO
Morinay J, Woodward BK, Russell AF, Sharp SP & Hatchwell BJ (2025) Ecological and demographic drivers of kin‐directed cooperation in a social bird: Insights from a long‐term study. Journal of Animal Ecology, 94(4), 485-500. View this article in WRRO
Laurenson K, Wood MJ, Birkhead TR, Priestley MDK, Sherley RB, Fayet AL, Guilford T, Hatchwell BJ & Votier SC (2025) Long‐term multi‐species demographic studies reveal divergent negative impacts of winter storms on seabird survival. Journal of Animal Ecology, 94(1), 139-153. View this article in WRRO
Halliwell C, Beckerman AP, Patrick SC & Hatchwell BJ (2024) Coordination of care reduces conflict and predation risk in a cooperatively breeding bird. Evolution Letters, 8(6), 764-773. View this article in WRRO
Borray-Escalante NA, Nuñez-Tobajas Z, Batllori X, Santos D, Clavell J, Domènech J, Arroyo L, Uribe F, Rodriguez-Pastor R, Pascual J , Carrillo-Ortiz JG et al (2024) Rewinding the invasion history of monk parakeets in Barcelona city: 1976-2022. Animal Biodiversity and Conservation, 47(1), 101-111. View this article in WRRO
Dawson Pell FSE, Hatchwell BJ, Ortega-Segalerva A & Senar JC (2024) Social associations are predicted by nest proximity but not kinship in a free-living social parrot. Biological Journal of the Linnean Society, 143(3). View this article in WRRO
Dawson Pell FSE, Senar JC, Ortega-Segalerva A & Hatchwell BJ (2024) Cooperation and conflict in the building and maintenance of the compound nests of monk parakeets Myiopsitta monachus. Animal Biodiversity and Conservation, 47(1), 9-17. View this article in WRRO
Lewis TC, Vargas IG, Vredenbregt C, Jimenez M, Hatchwell B, Beckerman AP & Childs DZ (2024) Nest-site selection and reproductive success of a critically endangered parrot, the Great Green Macaw (Ara ambiguus), in an anthropogenic landscape. Ibis, 166(2), 518-533. View this article in WRRO
Dawson Pell FSE, Hatchwell BJ, Carrillo-Ortiz JG, Ortega-Segalerva A, Uribe F & Senar JC (2023) Dispersal behaviour and settlement in an invasive bird: dispersers prefer their natal habitat. Animal Behaviour, 205, 139-148. View this article in WRRO
McCully FR, Descamps S, Harris SM, Mckendrick F, Gillies N, Cornell SJ, Hatchwell BJ & Patrick SC (2023) Links between personality, reproductive success and re‐pairing patterns in a long‐lived seabird. Ethology, 129(12), 686-700. View this article in WRRO
Halliwell C, Biddiscombe SJ & Hatchwell BJ (2023) Experimental variation of perceived predation risk does not affect coordination of parental care in long-tailed tits. Behavioral Ecology and Sociobiology, 77(9). View this article in WRRO
Halliwell C, Beckerman AP, Biddiscombe SJ, Germain M & Hatchwell BJ (2023) Coordination of care is facilitated by delayed feeding and collective arrivals in the long-tailed tit. Animal Behaviour, 201, 23-44. View this article in WRRO
Dawson Pell FSE, Senar JC & Hatchwell BJ (2023) Heterospecific nest material kleptoparasitism: observations of grey herons, Ardea cinerea, removing material from the nests of monk parakeets, Myiopsitta monachus. Arxius de Miscel·lània Zoològica, 21, 13-17. View this article in WRRO
Borray-Escalante NA, Baucells J, Carrillo-Ortiz JG, Hatchwell BJ & Senar JC (2023) Long distance dispersal of monk parakeets. Animal Biodiversity and Conservation, 46(1), 71-78. View this article in WRRO
Khwaja N, Preston S, Hatchwell B & Briskie J (2023) Recruitment, survival and breeding success in a declining rifleman population. New Zealand Journal of Ecology, 47(1).
McCully FR, Weimerskirch H, Cornell SJ, Hatchwell BJ, Cairo M & Patrick SC (2022) Partner intrinsic characteristics influence foraging trip duration, but not coordination of care in wandering albatrosses Diomedea exulans. Ecology and Evolution, 12(12).
Sturrock NJ, Hatchwell BJ, Firth JA & Green JP (2022) Who to help? Helping decisions in a cooperatively breeding bird with redirected care. Behavioral Ecology and Sociobiology, 76(6).
Halliwell C, Beckerman AP, Germain M, Patrick SC, Leedale AE & Hatchwell BJ (2022) Coordination of care by breeders and helpers in the cooperatively breeding long-tailed tit. Behavioral Ecology, 33(4), 844-858. View this article in WRRO
Roper M, Sturrock NJ, Hatchwell BJ & Green JP (2022) Individual variation explains ageing patterns in a cooperatively breeding bird, the long-tailed tit Aegithalos caudatus. Journal of Animal Ecology, 91(7), 1365-2656. View this article in WRRO
Senar JC, Moyà A, Pujol J, Tomas X & Hatchwell BJ (2021) Sex and age effects on monk parakeet home-range variation in the urban habitat. Diversity, 13(12).
Dawson Pell FSE, Senar JC, Franks DW & Hatchwell BJ (2021) Fine‐scale genetic structure reflects limited and coordinated dispersal in the colonial monk parakeet, Myiopsitta monachus. Molecular Ecology, 30(6), 1531-1544. View this article in WRRO
Ellison N, Hatchwell B, Biddiscombe S, Napper C & Potts J (2020) Mechanistic home range analysis reveals drivers of space use patterns for a non-territorial passerine. Journal of Animal Ecology, 89(12), 2763-2776. View this article in WRRO
Higgott CG, Evans KL & Hatchwell BJ (2020) Incubation in a temperate passerine: do environmental conditions affect incubation period duration and hatching success?. Frontiers in Ecology and Evolution, 8. View this article in WRRO
Leedale AE, Simeoni M, Sharp SP, Green JP, Slate J, Lachlan RF, Robinson EJH & Hatchwell BJ (2020) Cost, risk, and avoidance of inbreeding in a cooperatively breeding bird. Proceedings of the National Academy of Sciences, 117(27), 15724-15730.
Leedale AE, Lachlan RF, Robinson EJH & Hatchwell BJ (2020) Helping decisions and kin recognition in long-tailed tits : is call similarity used to direct help towards kin?. Philosophical Transactions of the Royal Society B: Biological Sciences, 375(1802). View this article in WRRO
Leedale AE, Li J & Hatchwell BJ (2020) Kith or kin? Familiarity as a cue to kinship in social birds. Frontiers in Ecology and Evolution, 8.
Dawson Pell FSE, Hatchwell BJ, Ortega-Segalerva A, Dawson DA, Horsburgh GJ & Senar JC (2020) Microsatellite characterisation and sex-typing in two invasive parakeet species, the monk parakeet Myiopsitta monachus and ring-necked parakeet Psittacula krameri. Molecular Biology Reports, 47(2), 1543-1550. View this article in WRRO
Li J, Lv L, Wang P, Wang Y, Hatchwell BJ & Zhang Z (2019) Sex-biased dispersal patterns of a social passerine: complementary approaches and evidence for a role of spatial scale. Biological Journal of the Linnean Society, 128(3), 592-602.
Li J, Wang Y, Lv L, Wang P, Hatchwell BJ & Zhang Z (2019) Context-dependent strategies of food allocation among offspring in a facultative cooperative breeder. Behavioral Ecology, 30(4), 975-985. View this article in WRRO
Khwaja N, Briskie JV & Hatchwell BJ (2019) Individuality, kin similarity and experimental playback of contact calls in cooperatively breeding riflemen. New Zealand Journal of Zoology, 46(4), 334-347. View this article in WRRO
Senar JC, Carrillo-Ortiz JG, Ortega-Segalerva A, Pell FSED, Pascual J, Arroyo L, Mazzoni D, Montalvo T & Hatchwell BJ (2019) The reproductive capacity of Monk Parakeets Myiopsitta monachus is higher in their invasive range. Bird Study, 66(1), 136-140. View this article in WRRO
Green J & Hatchwell BJ (2018) Inclusive fitness consequences of dispersal decisions in a cooperatively breeding bird, the long-tailed tit (Aegithalos caudatus). Proceedings of the National Academy of Sciences, 115(47), 12011-12016. View this article in WRRO
Leedale A, Sharp SP, Simeoni M, Robinson EJH & Hatchwell BJ (2018) Fine‐scale genetic structure and helping decisions in a cooperatively breeding bird . Molecular Ecology, 27(7), 1714-1726. View this article in WRRO
Khwaja N, Preston SAJ, Briskie JV & Hatchwell BJ (2018) Testing the predictions of sex allocation hypotheses in dimorphic, cooperatively breeding riflemen. Ecology and Evolution, 8(7), 3693-3701. View this article in WRRO
Khwaja N, Hatchwell BJ, Freckleton RP & Green JP (2017) Sex Allocation Patterns across Cooperatively Breeding Birds Do Not Support Predictions of the Repayment Hypothesis . American Naturalist, 190(4), 547-556. View this article in WRRO
Hatchwell BJ (2017) Replication in behavioural ecology: a comment on Ihle et al.. Behavioral Ecology, 28(2), 360. View this article in WRRO
Khwaja N, Preston SAJ, Hatchwell BJ, Briskie JV, Winney IS & Savage JL (2017) Flexibility but no coordination of visits in provisioning riflemen. Animal Behaviour, 125, 25-31.
Napper CJ & Hatchwell BJ (2016) Social dynamics in nonbreeding flocks of a cooperatively breeding bird: causes and consequences of kin associations. Animal Behaviour, 122, 23-35. View this article in WRRO
Bebbington K & Hatchwell BJ (2016) Coordinated parental provisioning is related to feeding rate and reproductive success in a songbird. Behavioral Ecology, 27(2), 652-659.
Green JP, Freckleton RP & Hatchwell BJ (2016) Variation in helper effort among cooperatively breeding bird species is consistent with Hamilton’s Rule. Nature Communications, 7(-), ---.
Sæther B-E, Grøtan V, Engen S, Coulson T, Grant PR, Visser ME, Brommer JE, Rosemary Grant B, Gustafsson L, Hatchwell BJ , Jerstad K et al (2016) Demographic routes to variability and regulation in bird populations. Nature Communications, 7(-), ---.
Preston SAJ, Briskie JV & Hatchwell BJ (2016) Adult helpers increase the recruitment of closely related offspring in the cooperatively breeding rifleman. Behavioral Ecology, 27(6), 1617-1626. View this article in WRRO
Acker P, Gregoire A, Rat M, Spottiswoode CN, van Dijk RE, Paquet M, Kaden JC, Pradel R, Hatchwell BJ, Covas R & Doutrelant C (2015) Disruptive viability selection on a black plumage trait associated with dominance. Journal of Evolutionary Biology, 28(11), 2027-2041. View this article in WRRO
van Dijk RE, Covas R, Doutrelant C, Spottiswoode CN & Hatchwell BJ (2015) Fine-scale genetic structure reflects sex-specific dispersal strategies in a population of sociable weavers (Philetairus socius).. Molecular Ecology. View this article in WRRO
Adams MJ, Robinson MR, Mannarelli M-E & Hatchwell BJ (2015) Social genetic and social environment effects on parental and helper care in a cooperatively breeding bird. Proceedings of the Royal Society B: Biological Sciences, 282(1810).
Crane JMS, Pick JL, Tribe AJ, Vincze E, Hatchwell BJ & Russell AF (2015) Chestnut-crowned babblers show affinity for calls of removed group members: a dual playback without expectancy violation. Animal Behaviour, 104, 51-57.
van Dijk RE, Kaden JC, Arguelles Tico A, Dawson DA, Burke T & Hatchwell BJ (2014) Cooperative investment in public goods is kin directed in communal nests of social birds. Ecology Letters, 17(9), 1141-1148.
Hatchwell BJ, Gullett PR & Adams MJ (2014) Helping in cooperatively breeding long-tailed tits: a test of Hamilton's rule.. Philos Trans R Soc Lond B Biol Sci, 369(1642), 20130565.
Napper CJ, Simeoni M, Hatchwell BJ, Sharp SP & McGowan A (2013) Dominance, not kinship, determines individual position within the communal roosts of a cooperatively breeding bird. Behavioral Ecology and Sociobiology, 1-11.
Gullett P, Hatchwell BJ, Robinson RA & Evans KL (2013) Phenological indices of avian reproduction: cryptic shifts and prediction across large spatial and temporal scales. Ecology and Evolution, 3(7), 1864-1877.
Preston SAJ, Dawson DA, Horsburgh GJ & Hatchwell BJ (2013) Characterisation of microsatellite loci in the rifleman (Acanthisitta chloris, Acanthisittidae, AVES) and their predicted genome locations. Conservation Genetics Resources, 5(2), 555-560.
van Dijk RE, Eising CM, Merrill RM, Karadas F, Hatchwell B & Spottiswoode CN (2013) Maternal effects in the highly communal sociable weaver may exacerbate brood reduction and prepare offspring for a competitive social environment. Oecologia, 171(2), 379-389.
Meade J, Hatchwell BJ, Blanchard JL & Birkhead TR (2012) The population increase of common guillemots Uria aalge on Skomer Island is explained by intrinsic demographic properties. Journal of Avian Biology.
Evans KL, Newton J, Gaston KJ, Sharp SP, Mcgowan A & Hatchwell BJ (2011) Colonisation of urban environments is associated with reduced migratory behaviour, facilitating divergence from ancestral populations. Oikos.
Lee J-W & Hatchwell BJ (2011) Do females adjust brood sex ratio according to males' genetic structures in a gregarious passerine, the vinous-throated parrotbill Paradoxornis webbianus?. Journal of Avian Biology.
Nam K-B, Meade J & Hatchwell BJ (2011) Do parents and helpers adjust their provisioning effort in relation to nestling sex in a cooperatively breeding bird?. Animal Behaviour.
Nam KB, Meade J & Hatchwell BJ (2011) Do parents and helpers adjust their provisioning effort in relation to nestling sex in a cooperatively breeding bird?. Animal Behaviour, 82(2), 303-309.
Beckerman AP, Sharp SP & Hatchwell BJ (2011) Predation and kin-structured populations: An empirical perspective on the evolution of cooperation. Behavioral Ecology, 22(6), 1294-1303.
Meade J, Nam K-B, Lee J-W & Hatchwell BJ (2011) An experimental test of the information model for negotiation of biparental care.. PLoS One, 6(5), e19684.
Sharp SP, Simeoni M, McGowan A, Nam KB & Hatchwell BJ (2011) Patterns of recruitment, relatedness and cooperative breeding in two populations of long-tailed tits. Animal Behaviour, 81(4), 843-849.
Sharp SP, Simeoni M, McGowan A, Nam K-B & Hatchwell BJ (2011) Patterns of recruitment, relatedness and cooperative breeding in two populations of long-tailed tits. Animal Behaviour.
Abbot P, Abe J, Alcock J, Alizon S, Alpedrinha JA, Andersson M, Andre JB, van Baalen M, Balloux F, Balshine S , Barton N et al (2011) Inclusive fitness theory and eusociality.. Nature, 471(7339), E1-E4.
Nam K-B, Meade J & Hatchwell BJ (2011) Brood sex ratio variation in a cooperatively breeding bird.. J Evol Biol, 24(4), 904-913.
Evans KL, Chamberlain DE, Hatchwell BJ, Gregory RD & Gaston KJ (2011) What makes an urban bird?. Global Change Biology, 17(1), 32-44.
Nakagawa S, Ockendon N, Gillespie DOS, Hatchwell BJ & Burke T (2011) Corrections for "Assessing the function of house sparrows' bib size using a flexible meta-analysis method [Behav Ecol 18: 831-840]". Behavioral Ecology, 22(2), 445-446.
Meade J & Hatchwell BJ (2010) No direct fitness benefits of helping in a cooperative breeder despite higher survival of helpers. BEHAV ECOL, 21(6), 1186-1194.
Evans KL, Hatchwell BJ, Parnell M & Gaston KJ (2010) A conceptual framework for the colonisation of urban areas: the blackbird Turdus merula as a case study.. Biol Rev Camb Philos Soc, 85(3), 643-667.
Nam K-B, Simeoni M, Sharp SP & Hatchwell BJ (2010) Kinship affects investment by helpers in a cooperatively breeding bird.. Proc Biol Sci, 277(1698), 3299-3306.
Lee JW, Lee YK & Hatchwell BJ (2010) Natal dispersal and philopatry in a group-living but noncooperative passerine bird, the vinous-throated parrotbill. Animal Behaviour, 79(5), 1017-1023.
Meade J, Nam K-B, Beckerman AP & Hatchwell BJ (2010) Consequences of 'load-lightening' for future indirect fitness gains by helpers in a cooperatively breeding bird.. J Anim Ecol, 79(3), 529-537.
Hatchwell BJ (2010) Cryptic Kin Selection: Kin Structure in Vertebrate Populations and Opportunities for Kin-Directed Cooperation. ETHOLOGY, 116(3), 203-216.
Lee JW, Kim HY & Hatchwell BJ (2010) Parental provisioning behaviour in a flock-living passerine, the Vinous-throated Parrotbill Paradoxornis webbianus. J ORNITHOL, 151(2), 483-490.
Lee J-W, Simeoni M, Burke T & Hatchwell BJ (2010) The consequences of winter flock demography for genetic structure and inbreeding risk in vinous-throated parrotbills, Paradoxornis webbianus.. Heredity (Edinb), 104(5), 472-481.
Hatchwell BJ (2009) The evolution of cooperative breeding in birds: kinship, dispersal and life history.. Philos Trans R Soc Lond B Biol Sci, 364(1533), 3217-3227.
Votier SC, Hatchwell BJ, Mears M & Birkhead TR (2009) Changes in the timing of egg-laying of a colonial seabird in relation to population size and environmental conditions. MAR ECOL-PROG SER, 393, 225-233.
Simeoni M, Dawson DA, Gentle LK, Coiffait L, Wolff K, Evans KL, Gaston KJ & Hatchwell BJ (2009) Characterization of 38 microsatellite loci in the European blackbird, Turdus merula (Turdidae, AVES). MOL ECOL RESOUR, 9(6), 1520-1526.
Lee JW, Kim MS, Burke T & Hatchwell BJ (2009) Extrapair paternity in a flock-living passerine, the vinous-throated parrotbill Paradoxornis webbianus. J AVIAN BIOL, 40(5), 469-474.
Evans KL, Gaston KJ, Frantz AC, Simeoni M, Sharp SP, McGowan A, Dawson DA, Walasz K, Partecke J, Burke T & Hatchwell BJ (2009) Independent colonization of multiple urban centres by a formerly forest specialist bird species.. Proc Biol Sci, 276(1666), 2403-2410.
Lee J-W, Jang B-S, Dawson DA, Burke T & Hatchwell BJ (2009) Fine-scale genetic structure and its consequence in breeding aggregations of a passerine bird.. Mol Ecol, 18(12), 2728-2739.
Evans KL, Gaston KJ, Sharp SP, McGowan A, Simeoni M & Hatchwell BJ (2009) Effects of urbanisation on disease prevalence and age structure in blackbird Turdus merula populations. OIKOS, 118(5), 774-782.
Hatchwell BJ, Sharp SP, Simeoni M & McGowan A (2009) Factors influencing overnight loss of body mass in the communal roosts of a social bird. FUNCT ECOL, 23(2), 367-372.
Evans KL, Gaston KJ, Sharp SP, McGowan A & Hatchwell BJ (2009) The effect of urbanisation on avian morphology and latitudinal gradients in body size. OIKOS, 118(2), 251-259.
Chamberlain DE, Cannon AR, Toms MP, Leech DI, Hatchwell BJ & Gaston KJ (2009) Avian productivity in urban landscapes: a review and meta-analysis. IBIS, 151(1), 1-18.
Nakagawa S, Lee J-W, Woodward BK, Hatchwell BJ & Burke T (2008) Differential selection according to the degree of cheating in a status signal.. Biol Lett, 4(6), 667-669.
Sharp SP, Baker MB, Hadfield JD, Simeoni M & Hatchwell BJ (2008) Natal dispersal and recruitment in a cooperatively breeding bird. OIKOS, 117(9), 1371-1379.
Votier SC, Birkhead TR, Oro D, Trinder M, Grantham MJ, Clark JA, McCleery RH & Hatchwell BJ (2008) Recruitment and survival of immature seabirds in relation to oil spills and climate variability.. J Anim Ecol, 77(5), 974-983.
Sharp SP, Simeoni M & Hatchwell BJ (2008) Dispersal of sibling coalitions promotes helping among immigrants in a cooperatively breeding bird.. Proc Biol Sci, 275(1647), 2125-2130.
Simeoni M, Dawson DA, Ross DJ, Chaline N, Burke T & Hatchwell BJ (2007) Characterization of 20 microsatellite loci in the long-tailed tit Aegithalos caudatus (Aegithalidae, AVES). MOL ECOL NOTES, 7(6), 1319-1322.
Hatchwell BJ (2007) Avian reproduction: role of ecology in the evolution of cooperative breeding.. Curr Biol, 17(19), R845-R847.
Nakagawa S, Gillespie DOS, Hatchwell BJ & Burke T (2007) Predictable males and unpredictable females: sex difference in repeatability of parental care in a wild bird population.. J Evol Biol, 20(5), 1674-1681.
Hatchwell BJ (2007) Historical perspectives and a focus on fitness.. Behav Processes, 76(2), 73-74.
Nakagawa S, Ockendon N, Gillespie DOS, Hatchwell BJ & Burke T (2007) Does the badge of status influence parental care and investment in house sparrows? An experimental test.. Oecologia, 153(3), 749-760.
Mcgowan A, Fowlie MK, Ross DJ & Hatchwell BJ (2007) Social organization of co-operatively breeding Long-tailed Tits Aegithalos caudatus: flock composition and kinship. IBIS, 149(1), 170-174.
Hatchwell BJ & Sharp SP (2006) Kin selection, constraints, and the evolution of cooperative breeding in long-tailed tits. ADV STUD BEHAV, 36, 355-395.
McGowan A, Sharp SP, Simeoni M & Hatchwell BJ (2006) Competing for position in the communal roosts of long-tailed tits. ANIM BEHAV, 72, 1035-1043.
Hatchwell BJ & Sharp SP (2006) Kin selection and recognition mechanisms in avian societies. J ORNITHOL, 147(5), 8-9.
Sharp SP & Hatchwell BJ (2006) Development of family specific contact calls in the Long-tailed Tit Aegithalos caudatus. IBIS, 148(4), 649-656.
Pizzari T, Birkhead TR, Blows MW, Brooks R, Buchanan KL, Clutton-Brock TH, Harvey PH, Hosken DJ, Jennions MD, Kokko H , Kotiaho JS et al (2006) Debating sexual selection and mating strategies.. Science, 312(5774), 689-697.
Lessells CM, Bennett ATD, Birkhead TR, Colegrave N, Dall SRX, Harvey PH, Hatchwell B, Hosken DJ, Hunt J, Moore AJ , Parker GA et al (2006) Debating sexual selection and mating strategies [2]. Science, 312(5774), 689-690.
Ghiselin MT (2006) Debating sexual selection and mating strategies.. Science, 312(5774), 689-697.
Votier SC, Hatchwell BJ, Beckerman A, McCleery RH, Hunter FM, Pellatt J, Trinder M & Birkhead TR (2005) Oil pollution and climate have wide-scale impacts on seabird demographics. ECOL LETT, 8(11), 1157-1164.
Cannon AR, Chamberlain DE, Toms MP, Hatchwell BJ & Gaston KJ (2005) Trends in the use of private gardens by wild birds in Great Britain 1995-2002. J APPL ECOL, 42(4), 659-671.
Sharp SP, McGowan A, Wood MJ & Hatchwell BJ (2005) Learned kin recognition cues in a social bird.. Nature, 434(7037), 1127-1130.
Sharp SP & Hatchwell BJ (2005) Individuality in the contact calls of cooperatively breeding long-tailed tits (Aegithalos caudatus). BEHAVIOUR, 142, 1559-1575.
Maccoll ADC & Hatchwell BJ (2004) Determinants of lifetime fitness in a cooperative breeder, the long-tailed tit Aegithalos caudatus. J ANIM ECOL, 73(6), 1137-1148.
McGowan A, Sharp SP & Hatchwell BJ (2004) The structure and function of nests of Long-Tailed Tits Aegithalos caudatus. FUNCT ECOL, 18(4), 578-583.
Hatchwell BJ, Russell AF, MacColl ADC, Ross DJ, Fowlie MK & McGowan A (2004) Helpers increase long-term but not short-term productivity in cooperatively breeding long-tailed tits. BEHAV ECOL, 15(1), 1-10.
MacColl ADC & Hatchwell BJ (2003) Heritability of parental effort in a passerine bird.. Evolution, 57(9), 2191-2195.
MacColl ADC & Hatchwell BJ (2003) HERITABILITY OF PARENTAL EFFORT IN A PASSERINE BIRD. Evolution, 57(9), 2191-2191.
McGowan A, Hatchwell BJ & Woodburn RJW (2003) The effect of helping behaviour on the survival of juvenile and adult long-tailed tits Aegithalos caudatus. J ANIM ECOL, 72(3), 491-499.
MacColl ADC & Hatchwell BJ (2003) Sharing of caring: nestling provisioning behaviour of long-tailed tit, Aegithalos caudatus, parents and helpers. ANIM BEHAV, 66, 955-964.
Hatchwell BJ, Ross DJ, Chaline N, Fowlie MK & Burke T (2002) Parentage in the cooperative breeding system of long-tailed tits, Aegithalos caudatus. ANIM BEHAV, 64, 55-63.
MacColl ADC & Hatchwell BJ (2002) Temporal variation in fitness payoffs promotes cooperative breeding in long-tailed tits Aegithalos caudatus.. Am Nat, 160(2), 186-194.
Hatchwell BJ, Wood MJ, Anwar MA, Chamberlain DE & Perrins CM (2001) The haematozoan parasites of Common Blackbirds Turdus merula: associations with host condition. IBIS, 143(3), 420-426.
Hatchwell BJ, Anderson C, Ross DJ, Fowlie MK & Blackwell PG (2001) Social organization of cooperatively breeding long-tailed tits: kinship and spatial dynamics. J ANIM ECOL, 70(5), 820-830.
Russell AF & Hatchwell BJ (2001) Experimental evidence for kin-biased helping in a cooperatively breeding vertebrate.. Proc Biol Sci, 268(1481), 2169-2174.
Hatchwell BJ, Ross DJ, Fowlie MK & McGowan A (2001) Kin discrimination in cooperatively breeding long-tailed tits.. Proc Biol Sci, 268(1470), 885-890.
Birkhead TR, Hatchwell BJ, Lindner R, Blomqvist D, Pellatt EJ, Griffiths R & Lifjeld JT (2001) Extra-Pair Paternity in the Common Murre. The Condor, 103(1), 158-162.
Birkhead TR, Hatchwell BJ, Lindner R, Blomqvist D, Pellatt EJ, Griffiths R & Lifjeld JT (2001) Extra-pair paternity in the Common Murre. CONDOR, 103(1), 158-162.
Hatchwell BJ, Russell AF, Ross DJ & Fowlie MK (2000) Divorce in cooperatively breeding long-tailed tits: a consequence of inbreeding avoidance?. Proc Biol Sci, 267(1445), 813-819.
Hatchwell BJ, Wood MJ, Anwar M & Perrins CM (2000) The prevalence and ecology of the haematozoan parasites of European blackbirds, Turdus merula. Canadian Journal of Zoology, 78(4), 684-687.
Hatchwell BJ, Wood MJ, Anwar M & Perrins CM (2000) The prevalence and ecology of the haematozoan parasites of European blackbirds, Turdus merula. CAN J ZOOL, 78(4), 684-687.
Hatchwell BJ & Komdeur J (2000) Ecological constraints, life history traits and the evolution of cooperative breeding.. Anim Behav, 59(6), 1079-1086.
Chamberlain DE, Hatchwell BJ & Perrins CM (1999) Importance of feeding ecology to the reproductive success of Blackbirds Turdus merula nesting in rural habitats. IBIS, 141(3), 415-427.
Komdeur J & Hatchwell BJ (1999) Reply from J. Komdeur and B.J. Hatchwell.. Trends Ecol Evol, 14(11), 448.
Komdeur J & Hatchwell BJ (1999) Kin recognition: function and mechanism in avian societies.. Trends Ecol Evol, 14(6), 237-241.
Barnard C, Komdeur J & Hatchwell BJ (1999) Still having trouble with relatives [1] (multiple letters). Trends in Ecology and Evolution, 14(11), 448.
Hatchwell BJ (1999) Investment strategies of breeders in avian cooperative breeding systems. AM NAT, 154(2), 205-219.
Hatchwell BJ, Fowlie MK, Ross DJ & Russell AF (1999) Incubation behavior of Long-tailed Tits: Why do males provision incubating females?. CONDOR, 101(3), 681-686.
Hatchwell BJ, Russell AF, Fowlie MK & Ross DJ (1999) Reproductive success and nest-site selection in a cooperative breeder: Effect of experience and a direct benefit of helping. AUK, 116(2), 355-363.
Hatchwell BJ, Chamberlain DE & Perrins CM (1996) The reproductive success of blackbirds Turdus merula in relation to habitat structure and choice of nest site. IBIS, 138(2), 256-262.
Hatchwell BJ & Russell AF (1996) Provisioning rules in cooperatively breeding long-tailed tits Aegithalos caudatus: An experimental study. P ROY SOC LOND B BIO, 263(1366), 83-88.
Langmore NE, Davies NB, Hatchwell BJ & Hartley IR (1996) Female song attracts males in the alpine accentor Prunella collaris. P ROY SOC LOND B BIO, 263(1367), 141-146.
Hatchwell BJ, Chamberlain DE & Perrins CM (1996) The demography of blackbirds Turdus merula in rural habitats: Is farmland a sub-optimal habitat?. J APPL ECOL, 33(5), 1114-1124.
Davies NB, Hartley IR, Hatchwell BJ & Langmore NE (1996) Female control of copulations to maximize male help: A comparison of polygynandrous alpine accentors, Prunella collaris, and dunnocks, P-modularis. ANIM BEHAV, 51, 27-47.
Chamberlain DE, Hatchwell BJ & Perrins CM (1995) Spaced out nests and predators: An experiment to test the effects of habitat structure. J AVIAN BIOL, 26(4), 346-349.
DAVIES NB, HARTLEY IR, HATCHWELL BJ, DESROCHERS A, SKEER J & NEBEL D (1995) THE POLYGYNANDROUS MATING SYSTEM OF THE ALPINE ACCENTOR, PRUNELLA-COLLARIS .1. ECOLOGICAL CAUSES AND REPRODUCTIVE CONFLICTS. ANIM BEHAV, 49(3), 769-788.
HARTLEY IR, DAVIES NB, HATCHWELL BJ, DESROCHERS A, NEBEL D & BURKE T (1995) THE POLYGYNANDROUS MATING SYSTEM OF THE ALPINE ACCENTOR, PRUNELLA-COLLARIS .2. MULTIPLE PATERNITY AND PARENTAL EFFORT. ANIM BEHAV, 49(3), 789-803.
EVANS MR & HATCHWELL BJ (1993) NEW SLANTS ON ORNAMENT ASYMMETRY. P ROY SOC LOND B BIO, 251(1332), 171-177.
HATCHWELL B & DAVIES N (1992) An experimental study of mating competition in monogamous and polyandrous dunnocks, : I. Mate guarding and copulations. Animal Behaviour, 43, 595-609.
EVANS MR & HATCHWELL BJ (1992) AN EXPERIMENTAL-STUDY OF MALE ADORNMENT IN THE SCARLET-TUFTED MALACHITE SUNBIRD .2. THE ROLE OF THE ELONGATED TAIL IN MATE CHOICE AND EXPERIMENTAL-EVIDENCE FOR A HANDICAP. BEHAV ECOL SOCIOBIOL, 29(6), 421-427.
EVANS MR & HATCHWELL BJ (1992) AN EXPERIMENTAL-STUDY OF MALE ADORNMENT IN THE SCARLET-TUFTED MALACHITE SUNBIRD .1. THE ROLE OF PECTORAL TUFTS IN TERRITORIAL DEFENSE. BEHAV ECOL SOCIOBIOL, 29(6), 413-419.
DAVIES NB & HATCHWELL BJ (1992) THE VALUE OF MALE PARENTAL CARE AND ITS INFLUENCE ON REPRODUCTIVE ALLOCATION BY MALE AND FEMALE DUNNOCKS. J ANIM ECOL, 61(2), 259-272.
HATCHWELL BJ & DAVIES NB (1992) AN EXPERIMENTAL-STUDY OF MATING COMPETITION IN MONOGAMOUS AND POLYANDROUS DUNNOCKS, PRUNELLA-MODULARIS .1. MATE GUARDING AND COPULATIONS. ANIM BEHAV, 43(4), 595-609.
HATCHWELL BJ & DAVIES NB (1992) AN EXPERIMENTAL-STUDY OF MATING COMPETITION IN MONOGAMOUS AND POLYANDROUS DUNNOCKS, PRUNELLA-MODULARIS .2. INFLUENCE OF REMOVAL AND REPLACEMENT EXPERIMENTS ON MATING SYSTEMS. ANIM BEHAV, 43(4), 611-622.
DAVIES NB, HATCHWELL BJ, ROBSON T & BURKE T (1992) PATERNITY AND PARENTAL EFFORT IN DUNNOCKS PRUNELLA-MODULARIS - HOW GOOD ARE MALE CHICK-FEEDING RULES. ANIM BEHAV, 43(5), 729-745.
WILEY RH, HATCHWELL BJ & DAVIES NB (1991) RECOGNITION OF INDIVIDUAL MALES SONGS BY FEMALE DUNNOCKS - A MECHANISM INCREASING THE NUMBER OF COPULATORY PARTNERS AND REPRODUCTIVE SUCCESS. ETHOLOGY, 88(2), 145-153.
BIRKHEAD TR, HATCHWELL BJ & DAVIES NB (1991) SPERM COMPETITION AND THE REPRODUCTIVE-ORGANS OF THE MALE AND FEMALE DUNNOCK PRUNELLA-MODULARIS. IBIS, 133(3), 306-311.
HATCHWELL BJ (1991) THE FEEDING ECOLOGY OF YOUNG GUILLEMOTS URIA-AALGE ON SKOMER ISLAND, WALES. IBIS, 133(2), 153-161.
HATCHWELL BJ (1991) AN EXPERIMENTAL-STUDY OF THE EFFECTS OF TIMING OF BREEDING ON THE REPRODUCTIVE SUCCESS OF COMMON GUILLEMOTS (URIA-AALGE). J ANIM ECOL, 60(3), 721-736.
HATCHWELL BJ & BIRKHEAD TR (1991) POPULATION-DYNAMICS OF COMMON GUILLEMOTS URIA-AALGE ON SKOMER ISLAND, WALES. ORNIS SCAND, 22(1), 55-59.
HATCHWELL BJ & PELLATT J (1990) INTRASPECIFIC VARIATION IN EGG COMPOSITION AND YOLK FORMATION IN THE COMMON GUILLEMOT (URIA-AALGE). J ZOOL, 220, 279-286.
HATCHWELL BJ & DAVIES NB (1990) PROVISIONING OF NESTLINGS BY DUNNOCKS, PRUNELLA-MODULARIS, IN PAIRS AND TRIOS - COMPENSATION REACTIONS BY MALES AND FEMALES. BEHAV ECOL SOCIOBIOL, 27(3), 199-209.
BURKE T, DAVIES NB, BRUFORD MW & HATCHWELL BJ (1989) PARENTAL CARE AND MATING-BEHAVIOR OF POLYANDROUS DUNNOCKS PRUNELLA-MODULARIS RELATED TO PATERNITY BY DNA FINGERPRINTING. NATURE, 338(6212), 249-251.
HATCHWELL BJ (1988) INTRASPECIFIC VARIATION IN EXTRA-PAIR COPULATION AND MATE DEFENSE IN COMMON GUILLEMOTS URIA AALGE. BEHAVIOUR, 107, 157-185.
Nakagawa S, Ockendon N, Gillespie DOS, Hatchwell BJ & Burke T () Assessing the function of house sparrows' bib size using a flexible meta-analysis method. Behavioral Ecology, 18(5), 831-840.

Book chapters

Cockburn A, Hatchwell BJ & Koenig WD (2017) Sociality in Birds, Comparative Social Evolution (pp. 320-353). Cambridge University Press
Hatchwell BJ (2016) Long-tailed tits: Ecological causes and fitness consequences of redirected helping, Cooperative Breeding in Vertebrates (pp. 39-57). Cambridge University Press
Komdeur J, Richardson DS & Hatchwell B (2008) Kin-Recognition Mechanisms in Cooperative Breeding Systems: Ecological Causes and Behavioral Consequences of Variation, Ecology of Social Evolution (pp. 175-193). Springer Berlin Heidelberg
Ekman J, Dickinson JL, Hatchwell BJ & Griesser M (2004) Delayed dispersal, Ecology and Evolution of Cooperative Breeding in Birds (pp. 35-47). Cambridge University Press
Dickinson JL & Hatchwell BJ (2004) Fitness consequences of helping, Ecology and Evolution of Cooperative Breeding in Birds (pp. 48-66). Cambridge University Press

Conference proceedings

Adams M, Robinson M & Hatchwell B (2013) Their genes are your environment: Social genetics of breeding behavior in birds. BEHAVIOR GENETICS, Vol. 43(6) (pp 505-505)

Preprints

Lewis TC, Gutiérrez Vargas I, Vredenbregt C, Jimenez M, Hatchwell B, Beckerman AP & Childs DZ (2023) Nest site selection and productivity of a critically endangered parrot, the great green macaw (Ara ambiguus), in an anthropogenic landscape, Cold Spring Harbor Laboratory.

Research group: Dr Alexandra Jebb de Calais
UKRI-funded Post-doctoral Research Associate (2023-2027). Constraints on adaptation in social animals: kin recognition mechanisms and the fitness consequences of discrimination rules. PI - BJH.
Dr Jennifer Morinay
UKRI-funded Post-doctoral Research Associate (2023-2027). Constraints on adaptation in social animals: kin recognition mechanisms and the fitness consequences of discrimination rules. PI - BJH.
Sarah Biddiscombe
UKRI-funded Technician (2022-2027). Constraints on adaptation in social animals: kin recognition mechanisms and the fitness consequences of discrimination rules. PI - BJH.
Dr Mickaël Jacquier
UKRI-funded Technician (2023-2027). Constraints on adaptation in social animals: kin recognition mechanisms and the fitness consequences of discrimination rules. PI - BJH.
Luke Nelson
UKRI-funded Technician (2023-2027). Constraints on adaptation in social animals: kin recognition mechanisms and the fitness consequences of discrimination rules. PI - BJH.
Tom Lewis
UKRI-funded Research Assistant (2023). Constraints on adaptation in social animals: kin recognition mechanisms and the fitness consequences of discrimination rules. PI - BJH.
Chay Halliwell
NERC-funded ACCE2 PhD student (2019-2023). Cooperation, conflict and the coordination of care in a social bird. Lead supervisor - BJH, co-supervisors - Dr Sam Patrick (University of Liverpool) and Dr Andrew Beckerman.
Fionnuala McCully
NERC-funded ACCE2 PhD student (2019-2023). How do Arctic seabirds coordinate care in a changing climate? Lead supervisor – Dr Sam Patrick (University of Liverpool), co-supervisor - BJH.
Natalia Borray
Independently funded PhD student (2022-2025). Population genetic structure and contemporary evolution in monk parakeets. Lead supervisor – Dr Juan Carlos Senar (Natural Sciences Museum of Barcelona; University of Barcelona), co-supervisor - BJH.
Joey Baxter
UKRI-funded PhD student (2023-2027). Are olfactory cues used to recognise kin in social birds? Lead-supervisor – BJH, co-supervisor – Dr Chris Cooney.

Teaching activities: I have been fascinated by natural history since I was young and becoming a behavioural ecologist was a natural progression from that early obsession. In common with my colleagues in Biosciences, my teaching is research-led, with the ambition of communicating my enthusiasm and knowledge about animal behaviour to students.

At Level 2, I teach on a field course in southern France, including guided exercises and independent project work in a stunning Mediterranean landscape. At Level 3, I coordinate Cooperation & Conflict (APS357, 10 credits), and I also run and/or teach on field courses to Portugal and Borneo (APS356, 20 credits). The aim of APS357 is to demonstrate how theory provides the framework that allows us to make sense of the extraordinarily diverse social behaviour that exists in the natural world, including our own behaviour. Field courses are highlights of the teaching year because of the quality of the projects conducted by students, and more importantly, because the opportunity to engage in research offered by such courses is often a critical moment in students’ development as scientists. At Level 4, I usually supervise two MBiolSci students per year. They are embedded within my research group, working alongside my PhD students, post-docs and other researchers on novel questions relating to some aspect of the behaviour and ecology of birds, involving field observations, experiments and analysis of long-term data. I also supervise MRes students in their research projects, typically one student per year.

Professional activities and memberships

International Society for Behavioral Ecology: President-elect (2014-2016), President (2016-2018) and Past-President (2018-2020)
External examiner: MSc in Applied Ecology, University of Exeter (Falmouth)
Editor: Behavioral Ecology (2010-2013)
Editor: Animal Behaviour (2005-2008)
Member of Council: Association for the Study of Animal Behaviour (2006-2009)
Member of Council: British Ornithologists’ Union (1999-2003)
Leverhulme Research Fellowship (2008-2009)